Pteranodontia







































Pteranodontians
Temporal range:
Early Cretaceous - Late Cretaceous, 127–66 Ma

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London Pteranodon.jpg
Mounted skeleton Pteranodon longiceps

Dinosaurs at CMNH 49.JPG
Fossil specimen of Nyctosaurus gracilis

Scientific classification e
Kingdom:
Animalia
Phylum:
Chordata
Order:
†Pterosauria
Suborder:
†Pterodactyloidea

Clade:
†Ornithocheiroidea

Clade:
†Pteranodontia
Marsh, 1887
Subgroups



  • †Nyctosauridae


  • †Pteranodontoidea


    • †Ornithocheiromorpha


    • †Pteranodontidae





Synonyms

Euornithocheira Unwin, 2003



Pteranodontia is an extinct group of ornithocheiroid pterosaurs that lived from the Early Cretaceous to the Late Cretaceous (middle Barremian to middle Campanian stages) of Asia, Europe, North America and South America.[1]




Contents






  • 1 Locomotion


  • 2 Classification


  • 3 "Wyomingopteryx"


  • 4 References





Locomotion


Pteranodontians, like other pterosaurs, are considered to have been skilled fliers as well as adept at moving on the ground. Evidence from footprints shows that most pterosaurs did not sprawl their limbs to a large degree, as in modern reptiles, but rather held the limbs relatively erect when walking, like dinosaurs. While no pteranodontian footprints are known, it is likely that they also walked erect.[2] Among pterosaurs, pteranodontians had unusually uneven limb proportions, with the forelimbs much longer than the hind limbs. This would likely have required them to use unique modes of locomotion when on the ground compared to other pterosaurs. It is possible that pteranodontians ran (but not walked) bipedally, or that they used a hopping gait.[2] Pterosaur researcher Mike Habib has noted that the limbs proportions of pteranodontians like Anhanguera are consistent with hopping.[3]


Pteranodontians were among the last of the world's pterosaur faunas. The species Piksi barbarulna and a few potential pteranodontids and nyctosaurs have all been found dating from the Campanian to the Maastrichtian ages of the Late Cretaceous period.[4]



Classification


Pteranodontia was originally named by O.C. Marsh in 1876. In 2003, it was given a phylogenetic definition by David Unwin as the common ancestor of Pteranodon and Nyctosaurus plus all its descendants. Though Marsh had originally named this group based on the shared absence of teeth in those species, most analyses show that all of the traditional "ornithocheiroid" pterosaurs are also members of this clade.


Below is a cladogram showing the results of a phylogenetic analysis first presented by Andres and Myers in 2014, and updated with additional data by Andres and colleagues in 2018.[5].mw-parser-output table.clade{border-spacing:0;margin:0;font-size:100%;line-height:100%;border-collapse:separate;width:auto}.mw-parser-output table.clade table.clade{width:100%}.mw-parser-output table.clade td{border:0;padding:0;vertical-align:middle;text-align:center}.mw-parser-output table.clade td.clade-label{width:0.8em;border:0;padding:0 0.2em;vertical-align:bottom;text-align:center}.mw-parser-output table.clade td.clade-slabel{border:0;padding:0 0.2em;vertical-align:top;text-align:center}.mw-parser-output table.clade td.clade-bar{vertical-align:middle;text-align:left;padding:0 0.5em}.mw-parser-output table.clade td.clade-leaf{border:0;padding:0;text-align:left;vertical-align:middle}.mw-parser-output table.clade td.clade-leafR{border:0;padding:0;text-align:right}









Ornithocheiroidea












 

Azhdarchoidea
 
 Pteranodontoidea 












 

Ornithocheiromorpha
 
 Pteranodontia 












 








Pteranodontidae












 












 

Pteranodon sternbergi
 
 

Pteranodon longiceps
 
 
 

Tethydraco regalis
 
 
 

Nyctosauridae

















 

Alamodactylus byrdi
 
 

Volgadraco bogolubovi
 
 












 

Cretornis hlavaci
 
 












 












 

Alcione elanius
 
 

Simurghia robusta
 
 
 












 

Muzquizopteryx coahuilensis
 
 












 












 

Barbaridactylus grandis
 
 

Nyctosaurus lamegoi
 
 
 












 

Nyctosaurus nanus
 
 

Nyctosaurus gracilis
 
 
 
 
 
 
 
 
 
 


"Wyomingopteryx"


The name "Wyomingopteryx" appears in a painting of Morrison prehistoric animals by Robert Bakker. However, this binomen is a nomen nudum, and it is possible that Bakker may have intended to coin "Wyomingopteryx" for the Istiodactylus-like specimen TATE 5999 because that specimen is found in Wyoming.[6][7]



References





  1. ^ Richard J. Butler; Stephen L. Brusatte; Brian B. Andres; Roger B. J. Benson (2012). "How do geological sampling biases affect studies of morphological evolution in deep time? A case study of the Pterosauria (Reptilia: Archosauria)". Evolution. 66 (1): 147–162. doi:10.1111/j.1558-5646.2011.01415.x..mw-parser-output cite.citation{font-style:inherit}.mw-parser-output .citation q{quotes:"""""""'""'"}.mw-parser-output .citation .cs1-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/thumb/6/65/Lock-green.svg/9px-Lock-green.svg.png")no-repeat;background-position:right .1em center}.mw-parser-output .citation .cs1-lock-limited a,.mw-parser-output .citation .cs1-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/thumb/d/d6/Lock-gray-alt-2.svg/9px-Lock-gray-alt-2.svg.png")no-repeat;background-position:right .1em center}.mw-parser-output .citation .cs1-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/thumb/a/aa/Lock-red-alt-2.svg/9px-Lock-red-alt-2.svg.png")no-repeat;background-position:right .1em center}.mw-parser-output .cs1-subscription,.mw-parser-output .cs1-registration{color:#555}.mw-parser-output .cs1-subscription span,.mw-parser-output .cs1-registration span{border-bottom:1px dotted;cursor:help}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/thumb/4/4c/Wikisource-logo.svg/12px-Wikisource-logo.svg.png")no-repeat;background-position:right .1em center}.mw-parser-output code.cs1-code{color:inherit;background:inherit;border:inherit;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;font-size:100%}.mw-parser-output .cs1-visible-error{font-size:100%}.mw-parser-output .cs1-maint{display:none;color:#33aa33;margin-left:0.3em}.mw-parser-output .cs1-subscription,.mw-parser-output .cs1-registration,.mw-parser-output .cs1-format{font-size:95%}.mw-parser-output .cs1-kern-left,.mw-parser-output .cs1-kern-wl-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right,.mw-parser-output .cs1-kern-wl-right{padding-right:0.2em}


  2. ^ ab Witton, M.P.; Habib, M.B. (2010). "On the Size and Flight Diversity of Giant Pterosaurs, the Use of Birds as Pterosaur Analogues and Comments on Pterosaur Flightlessness". PLoS ONE. 5 (11): e13982. doi:10.1371/journal.pone.0013982. PMC 2981443. PMID 21085624.


  3. ^ Habib, M. (2011). "Dinosaur Revolution: Anhanguera." H2VP: Paleobiomechanics. Weblog entry, 20-SEP-2011. Accessed 28-SEP-2011: http://h2vp.blogspot.com/2011/09/dinosaur-revolution-anhanguera.html


  4. ^ Federico L. Agnolin & David Varricchio (2012). "Systematic reinterpretation of Piksi barbarulna Varricchio, 2002 from the Two Medicine Formation (Upper Cretaceous) of Western USA (Montana) as a pterosaur rather than a bird" (PDF). Geodiversitas. 34 (4): 883–894. doi:10.5252/g2012n4a10.


  5. ^ Andres, B.; Clark, J.; Xu, X. (2014). "The Earliest Pterodactyloid and the Origin of the Group". Current Biology. 24: 1011–6. doi:10.1016/j.cub.2014.03.030. PMID 24768054.


  6. ^ Bakker, R.T. (1994) Unearthing the Jurassic. In: Science Year 1995. World Book Inc.:Chicago, London, Sydney, Toronto, 76-89.
    ISBN 0-7166-0595-3.


  7. ^ Bakker, R.T. (1998). "Dinosaur mid-life crisis: the Jurassic-Cretaceous transition in Wyoming and Colorado". In Lucas, Spencer G.; Kirkland, James I.; Estep, J.W. Lower and Middle Cretaceous Terrestrial Ecosystems 14. New Mexico Museum of Natural History and Science Bulletin. pp. 67–77.













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