Graft-chimaera






The small tree + Laburnocytisus 'Adamii' is a spectacular example of a graft-chimaera


In horticulture, a graft-chimaera may arise in grafting at the point of contact between rootstock and scion and will have properties intermediate between those of its "parents". A graft-chimaera is not a true hybrid but a mixture of cells, each with the genotype of one of its "parents": it is a chimaera. Hence, the once widely used term "graft-hybrid" is not descriptive; it is now frowned upon.


Propagation is by cloning only. In practice graft-chimaeras are not noted for their stability and may easily revert to one of the "parents".




Contents






  • 1 Nomenclature


  • 2 In Darwin's works


  • 3 See also


  • 4 References





Nomenclature


Article 21 of the ICNCP stipulates that a graft-chimaera can be indicated either by


  • a formula: the names of both "parents", in alphabetical order, joined by the plus sign "+":


Crataegus + Mespilus

  • a name:

    • if the "parents" belong to different genera a name may be formed by joining part of one generic name to the whole of the other generic name. This name must not be identical to a generic name published under the International Code of Nomenclature for algae, fungi, and plants (ICN). For example + Crataegomespilus is the name for the graft-chimaera which may also be indicated by the formula Crataegus + Mespilus. This name is clearly different from ×Crataemespilus, the name under the ICN for the true hybrid between Crataegus and Mespilus, which can also be designated by the formula Crataegus × Mespilus.

    • if both "parents" belong to the same genus the graft-chimaera may be given a cultivar name. For example Syringa 'Correlata' is a graft-chimaera involving Syringa vulgaris (common lilac) and Syringa × chinensis (Rouen lilac, which is itself a hybrid between S. vulgaris and S. laciniata). No plus sign is used, because both "parents" belong to the genus Syringa.



A graft-chimaera cannot have a species name, because it is simultaneously two species. Although + Laburnocytisus 'Adamii', for example, is sometimes seen written as if it were a species (+ Laburnocytisus adamii), this is incorrect.



In Darwin's works


Charles Darwin «The Variation of Animals and Plants Under Domestication», 1868 г.:


I will therefore give all the facts which I have been able to collect on the formation of hybrids between distinct species or varieties, without the intervention of the sexual organs. For if, as I am now convinced, this is possible, it is a most important fact, which will sooner or later change the views held by physiologists with respect to sexual reproduction. A sufficient body of facts will afterwards be adduced, showing that the segregation or separation of the characters of the two parent-forms by bud-variation, as in the case of Cytisus adami, is not an unusual though a striking phenomenon. We shall further see that a whole bud may thus revert, or only half, or some smaller segment.



See also




  • Bizzaria - a very popular graft chimaera in Citrus.

  • Chimera (genetics)#Plants

  • Grafting



References










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