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Paleontology or palaeontology (from Greek: paleo, "ancient"; ontos, "being"; and logos, "knowledge") is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.^[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2012.

Note: In 2012 International Code of Zoological Nomenclature was amended, with new regulations allowing the publication of new names and nomenclatural acts in zoology after 2011 in works "produced in an edition containing simultaneously obtainable copies by a method that assures (...) widely accessible electronic copies with fixed content and layout", provided that the work is registered in ZooBank before it is published, the work itself states the date of publication with evidence that registration has occurred, and the ZooBank registration states both the name of an electronic archive intended to preserve the work and the ISSN or ISBN associated with the work.^[2]^[3] New scientific names appearing in electronic works are not required to be registered in ZooBank, only the works themselves are.^[2] Works containing descriptions of some of the taxa listed below weren't printed on paper in 2012; however, the taxa that were described in works which were registered in ZooBank in 2012 are listed as valid.

Plants

Gymnosperms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Anomozamites villosus^[4]	Sp. nov	Valid	Pott et al.	Probably late Middle Jurassic	Daohugou Beds	China	A member of Bennettitales. Originally described as a species of Anomozamites, subsequently transferred to the genus Wielandiella.^[5]^[6]
Calocedrus huashanensis^[7]	Sp. nov	Valid	Shi, Zhou, & Xie	Oligocene	Ningming Formation	China	An extinct species in the modern genus Calocedrus
Dacrycarpus puertae^[8]	Sp. nov	Valid	Wilf	Eocene		Argentina	A species of Dacrycarpus
Duartenia^[9]	Gen. et sp. nov	Valid	Mohr et al.	Early Cretaceous	Crato Formation	Brazil	A possible member of Cheirolepidiaceae. The type species is Duartenia araripensis.
Eathiestrobus^[10]	Gen. et sp. nov	Valid	Rothwell et al.	Late Jurassic (Kimmeridgian)	Kimmeridge Clay Formation	United Kingdom	A member of Pinaceae. The type species is Eathiestrobus mackenziei.
Eretmonia macloughlinii^[11]	Sp. nov	Valid	Ryberg, Taylor & Taylor	Late Permian	Upper Buckley Formation	Antarctica	A member of Glossopteridales.
Fokienia shengxianensis^[12]	Sp. nov	Valid	He, Sun & Liu	Late Miocene	Shengxian Formation Xiaolongtan Formation	China	Originally described as a species of Fokienia, but subsequently transferred to the genus Calocedrus.^[13]
Ginkgo cranei^[14]	Sp. nov	Valid	Zhou, Quan, & Liu	upper Paleocene	Sentinel Butte Formation, North Dakota	USA	An extinct species in the modern genus Ginkgo
Ginkgo gomolitzkyana^[15]	Sp. nov	Valid	Nosova	Middle Jurassic	Angren Formation	Uzbekistan	A member of Ginkgoales.
Ginkgo huolinhensis^[16]	Sp. nov	Valid	Dong & Sun	Early Cretaceous	Huolinhe Formation	China
Glenrosa falcata^[17]	Sp. nov	Valid	Gomez, Ewin & Daviero-Gomez	Early Cretaceous (Barremian)	Calizas de La Huergina Formation	Spain	A conifer.
Kuvakospermum^[18]	Gen. et sp. nov	Valid	Naugolnykh & Sidorov	Permian (Wordian)		Russia	A seed fern belonging to the family Peltaspermaceae. Genus includes new species K. pedatum.
Lidgettoniopsis^[19]	Gen. et sp. nov	Valid	Ryberg, Taylor & Taylor	Late Permian	Beacon Supergroup (upper Buckley Formation)	Antarctica	A seed fern belonging to the group Dictyopteridiales and the family Eretmoniaceae. Genus includes new species L. ramulus.
Picea burtonii^[20]	Sp. nov	Valid	Klymiuk & Stockey	Early Cretaceous (Valanginian)		Canada	A spruce.
Pinus yorkshirensis^[21]	Sp. nov	Valid	Ryberg et al.	Early Cretaceous (Hauterivian-Barremian)	Speeton Clay Formation	United Kingdom	A pine.
Plyophyllioxylon^[22]	Gen. et sp. nov	Valid	Feng et al.	Permian (Roadian/Wordian)	Lower Shihhotse Formation	China	A conifer of uncertain phylogenetic placement. Genus includes new species P. hulstaiense.
Rehezamites^[23]	Gen. et sp. nov	Valid	Pott et al.	Early Cretaceous (Aptian)	Yixian Formation	China	A seed plant of uncertain phylogenetic placement. Genus includes new species R. anisolobus.
Scutum leiophyllum^[19]	Sp. nov	Valid	Ryberg, Taylor & Taylor	Late Permian	Beacon Supergroup (upper Buckley Formation)	Antarctica	A member of Glossopteridales belonging to the family Dictyopteridiaceae.
Vetlugospermum^[24]	Gen. et sp. nov	Valid	Naugolnykh	Early Triassic		Russia	A seed fern belonging to the group Peltaspermales. Genus includes new species V. rombicum.

Angiosperms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Archaeampelos senonica^[25]	Sp. nov	Valid	Alekseev	Late Cretaceous		Russia
Baasia^[26]	Gen. et sp. nov	Valid	Estrada-Ruiz et al.	Late Cretaceous (probably early to middle Maastrichtian)	McRae Formation	United States	A member of the family Celastraceae. Genus includes new species B. armendarisense.
Bertilanthus^[27]	Gen. et sp. nov	Valid	Friis & Pedersen	Late Cretaceous		Sweden	An asterid. Genus includes new species B. scanicus.
Camptodromites sibiricus^[25]	Sp. nov	Valid	Alekseev	Late Cretaceous		Russia
Celastrinites lanceolatus^[25]	Sp. nov	Valid	Alekseev	Late Cretaceous		Russia
Chaneya hainanensis^[28]	Sp. nov	Valid	Feng & Jin	Eocene	Changchang Basin	China	A fossil fruit.
Cissites basicordatus^[25]	Sp. nov	Valid	Alekseev	Late Cretaceous		Russia
Cissus lombardii^[29]	Sp. nov	Valid	Manchester, Chen & Lott	Early Oligocene	Mancora Formation	Peru	A species of Cissus.
Eucalyptus caldericola^[30]	Sp. nov	Valid	Hermsen, Gandolfo & Zamaloa	Eocene (Ypresian)		Argentina	A species of Eucalyptus.
Eucalyptus frenguelliana^[30]	Sp. nov	Valid	Gandolfo & Zamaloa in Hermsen, Gandolfo & Zamaloa	Eocene (Ypresian)		Argentina	A species of Eucalyptus.
Eucalyptus lynchiae^[30]	Sp. nov	Valid	Gandolfo & Hermsen in Hermsen, Gandolfo & Zamaloa	Eocene (Ypresian)		Argentina	A species of Eucalyptus.
Fulleroxylon^[26]	Gen. et sp. nov	Valid	Estrada-Ruiz et al.	Late Cretaceous (probably early to middle Maastrichtian)	McRae Formation	United States	A member of the family Myrtaceae. Genus includes new species F. armendarisense.
Juglandiphyllites microdentatus^[25]	Sp. nov	Valid	Alekseev	Late Cretaceous		Russia
Laurinoxylon stichkai^[31]	Sp. nov	Valid	Boonchai & Manchester	Early Eocene	Bridger Formation	United States	A member of Lauraceae described on the basis of fossil wood.
Laurophyllum calicarioides^[32]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Litsea.
Laurophyllum lacustris^[32]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Beilschmiedia.
Laurophyllum maarensis^[32]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Cryptocarya.
Laurophyllum microphyllum^[32]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Cryptocarya.
Laurophyllum otagoensis^[32]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Beilschmiedia.
Laurophyllum sylvestris^[32]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Beilschmiedia.
Laurophyllum taieriensis^[32]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Cryptocarya.
Laurophyllum vulcanicola^[32]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Beilschmiedia.
Laurophyllum waipiata^[32]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Cryptocarya.
Litseopsis nova-zelandiae^[32]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae.
Mahonia grimmii^[33]	Sp. nov	Valid	Güner & Denk	Miocene	Eskihisar Formation	Turkey	A species of Mahonia.
Mahonia somaensis^[33]	Sp. nov	Valid	Güner & Denk	Miocene	Soma Formation	Turkey	A species of Mahonia.
Morinda chinensis^[34]	Sp. nov	Valid	Shi et al.	Eocene	Changchang Formation	China	A species of Morinda.
Newtonia mushensis^[35]	Sp. nov	Valid	Pan et al.	Early Miocene		Ethiopia	A species of Newtonia.
Palaeophytocrene hammenii^[36]	Sp. nov	Valid	Stull in Stull et al.	Middle-late Paleocene	Bogotá Formation	Colombia	A member of Icacinaceae belonging to the tribe Phytocreneae.
Palaeophytocrene piggae^[36]	Sp. nov	Valid	Stull in Stull et al.	Late Paleocene	Sentinel Butte Formation	United States	A member of Icacinaceae belonging to the tribe Phytocreneae.
Pygmaeoxylon^[26]	Gen. et sp. nov	Valid	Estrada-Ruiz et al.	Late Cretaceous (probably early to middle Maastrichtian)	McRae Formation	United States	A magnoliid of uncertain phylogenetic placement. Genus includes new species P. paucipora.
Pyrenacantha austroamericana^[36]	Sp. nov	Valid	Stull in Stull et al.	Late early Oligocene		Peru	A species of Pyrenacantha.
Schenkeriphyllum glanduliferum^[37]	Gen. et sp. nov	Valid	Mohr, Coiffard & Bernardes-de-Oliveira	Early Cretaceous (Aptian)	Crato Formation	Brazil	Fossil magnolialean angiosperm.
Wilsonoxylon^[31]	Gen. et sp. nov	Valid	Boonchai & Manchester	Early Eocene	Bridger Formation	United States	A member of Canellaceae described on the basis of fossil wood. Genus includes new species W. edenense.

Other plants

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Korallipteris^[38]	Gen. et comb. nov	Valid	Vera & Passalia	Triassic to Miocene		Antarctica Argentina Chile New Zealand^[39]	A fern of uncertain phylogenetic placement. A new genus for "Gleichenia" argentinica (Berry) emend. Herbst (1962); genus also includes several other species originally assigned to the genera Gleichenia, Gleichenites or Microphyllopteris.
Millerocaulis tekelili^[40]	Sp. nov	Valid	Vera	Early Cretaceous (Aptian)	Cerro Negro Formation	Antarctica (Livingston Island)	A member of the family Osmundaceae. Originally described as a species of Millerocaulis; transferred to the genus Claytosmunda by Bomfleur, Grimm & McLoughlin (2017).^[41]
Osmundopsis rafaelii^[42]	Sp. nov	Valid	Escapa & Cúneo	Early Jurassic		Argentina	A member of Osmundaceae.
Riccardiothallus^[43]	Gen. et sp. nov	Valid	Guo et al.	Devonian (Pragian)	Posongchong Formation	China	A liverwort belonging to the family Aneuraceae. Genus includes new species R. devonicus.
Todites cacereii^[42]	Sp. nov	Valid	Escapa & Cúneo	Early Jurassic		Argentina	A member of Osmundaceae.
Winslowia^[44]	Gen. et sp. nov	Valid	Dunn et al.	Carboniferous (Serpukhovian)	Pride Mountain Formation	United States	A member of Isoetales belonging to the group Chaloneriaceae. Genus includes new species W. tuscumbiana.

Arthropods

The following is a summary of the arthopods described in 2012

41 arachnids

105 crustaceans

362 insects

5 merostomatans

51 trilobites

9 other arthropods

Bryozoans

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Acupipora^[45]	Gen. et comb. nov	Valid	Gorjunova & Weiss	Late Carboniferous		Russia	A member of Fenestellida. The type species is "Polypora" subborealis Schulga-Nesterenko (1951).
Admirandopora^[46]	Nom. nov.	Valid	Ariunchimeg	Lower Carboniferous		Mongolia	A replacement name for the genus Admiranda Ariunchimeg, 1996.
Antoniettella^[47]	Gen. et sp. nov	Valid	Di Martino & Taylor	Early or middle Miocene		Indonesia	A cribrimorph ascophoran. The type species is Antoniettella exigua.
Argentinodictya^[48]	Gen. et sp. nov	Valid	Ernst & Carrera	Late Ordovician (Sandbian)		Argentina	A cryptostome bryozoan, a member of Ptilodictyina. The type species is Argentinodictya lenticulata.
Bigeyina ibera^[49]	Sp. nov.	Valid	Ernst	Devonian		Spain	A member of Fenestrata.
Celleporina medoborensis^[50]	Sp. nov	Valid	Hara & Jasionowski	Miocene (early Sarmatian)		Ukraine	A celleporid cheilostome bryozoan, a species of Celleporina.
Chasmatopora rossae^[48]	Sp. nov	Valid	Ernst & Carrera	Late Ordovician (Sandbian)		Argentina	A phyloporinine, a species of Chasmatopora.
Dissotrypa punctata^[49]	Sp. nov.	Valid	Ernst	Devonian		Spain	A member of Fenestrata.
Dissotrypa robusta^[49]	Sp. nov.	Valid	Ernst	Devonian		Spain	A member of Fenestrata.
Eosemicoscinium serratum^[49]	Sp. nov.	Valid	Ernst	Devonian		Spain	A member of Fenestrata.
Hemitrypa cantabrica^[49]	Sp. nov.	Valid	Ernst	Devonian		Spain	A member of Fenestrata.
Iberofenestella^[49]	Gen. et sp. nov.	Valid	Ernst	Devonian		Spain	A member of Fenestrata. Genus includes new species I. wolfae.
Lunostoma^[51]	Gen. et sp. nov	Valid	Ernst, Taylor, Bohatý & Wyse Jackson	Middle Devonian		Germany	A cryptostome bryozoan. The type species is Lunostoma pulchra.
Parvohallopora parvula^[48]	Sp. nov	Valid	Ernst & Carrera	Late Ordovician (Sandbian)		Argentina	A trepostome bryozoan, a species of Parvohallopora.
Paucipora akishinensis^[45]	Sp. nov	Valid	Gorjunova & Weiss	Late Carboniferous (Moscovian)	Domodedovo Formation	Russia	A member of Fenestellida.
Prolixicella ibera^[49]	Sp. nov.	Valid	Ernst	Devonian		Spain	A member of Fenestrata.
Prolixicella parva^[49]	Sp. nov.	Valid	Ernst	Devonian		Spain	A member of Fenestrata.
Ptilodictya intermedia^[48]	Sp. nov	Valid	Ernst & Carrera	Late Ordovician (Sandbian)		Argentina	A cryptostome bryozoan, a species of Ptilodictya.
Pyrisinella^[52]	Gen. et comb. nov	Valid	Di Martino & Taylor	Late Cretaceous (Maastrichtian)	Peedee Formation Prairie Bluff Chalk Ripley Formation	United States	A member of Cheilostomata belonging to the family Pyrisinellidae. The type species is "Setosinella" meniscacantha Taylor & McKinney (2006).
Rectifenestella elegantula^[49]	Sp. nov.	Valid	Ernst	Devonian		Spain	A member of Fenestrata.
Rectifenestella villayandrensis^[49]	Sp. nov.	Valid	Ernst	Devonian		Spain	A member of Fenestrata.
Rorypora^[53]	Gen. et comb. nov.	Valid	Taylor	Jurassic		Switzerland	A new genus for "Diastopora" retiformis.
Setosinella perfluxa^[52]	Sp. nov	Valid	Di Martino & Taylor	Miocene (Langhian)		Indonesia	A member of Cheilostomata belonging to the family Pyrisinellidae.
Spinisinella^[52]	Gen. et sp. nov	Valid	Di Martino & Taylor	Late Cretaceous (Cenomanian)		Czech Republic Germany^[54]	A member of Cheilostomata belonging to the family Pyrisinellidae. The type species is S. zagorseki.
Trigonodictya parvula^[48]	Sp. nov	Valid	Ernst & Carrera	Late Ordovician (Sandbian)		Argentina	A cryptostome bryozoan, a species of Trigonodictya.

Brachiopods

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Amydroptychus markowitzi^[55]	sp nov	Valid	Feldman et al.	Callovian	Mughanniyya Formation	Jordan	A tetrarhynchiid rhynchonellide, a species of Amydroptychus.
Anisopleurella tricostata^[56]	Sp. nov	Valid	Rasmussen, Harper & Blodgett	Late Ordovician	Farewell Terrane	United States	A member of Strophomenida belonging to the group Plectambonitoidea and the family Sowerbyellidae.
Bolilaspirifer riccardii^[57]	Sp. nov	Valid	Manceñido in Damborenea & Manceñido	Late Triassic		Argentina	A lepismatinid brachiopod, a species of Bolilaspirifer.
Burrirhynchia angustisinuata^[58]	sp nov	Valid	Smirnova	Early Cretaceous		Russia	A cyclothyridid rhynchonellide, a species of Burrirhynchia.
Burrirhynchia latimarginata^[58]	sp nov	Valid	Smirnova	Early Cretaceous		Russia	A cyclothyridid rhynchonellide, a species of Burrirhynchia.
Burrirhynchia oweni^[58]	sp nov	Valid	Smirnova	Early Cretaceous		Russia	A cyclothyridid rhynchonellide, a species of Burrirhynchia.
Burrirhynchia subgrasiana^[58]	sp nov	Valid	Smirnova	Early Cretaceous		Russia	A cyclothyridid rhynchonellide, a species of Burrirhynchia.
Callositella^[56]	Gen. et sp. nov	Valid	Rasmussen, Harper & Blodgett	Late Ordovician	Farewell Terrane	United States	A member of Orthida belonging to the group Enteletoidea and the family Saukrodictyidae. The type species is C. cheeneetnukensis.
Christiania aseptata^[56]	Sp. nov	Valid	Rasmussen, Harper & Blodgett	Late Ordovician	Farewell Terrane	United States	A member of Strophomenida belonging to the family Christianiidae.
Cincinnetina^[59]	Gen, sp. et comb. nov	Valid	Jin	Late Ordovician		United States	A dalmanellid brachiopod, with C. multisecta, C. meeki, C. minnesotensis
Craspedelia potterella^[56]	Sp. nov	Valid	Rasmussen, Harper & Blodgett	Late Ordovician	Farewell Terrane	United States	A member of Strophomenida belonging to the group Plectambonitoidea and the family Bimuriidae.
Crurithyris tazawai^[60]	sp nov	Valid	He et al..	Late Permian (Changhsingian)		China	An ambocoeliid brachiopod, a species of Crurithyris.
Cyclothyris aliformis^[58]	sp nov	Valid	Smirnova	Early Cretaceous		Russia	A cyclothyridid rhynchonellide, a species of Burrirhynchia.
Cyclothyris alikentica^[58]	sp nov	Valid	Smirnova	Early Cretaceous		Russia	A cyclothyridid rhynchonellide, a species of Burrirhynchia.
Cyclothyris burgemakensis^[58]	sp nov	Valid	Smirnova	Early Cretaceous		Russia	A cyclothyridid rhynchonellide, a species of Burrirhynchia.
Cyclothyris dagestanica^[58]	sp nov	Valid	Smirnova	Early Cretaceous		Russia	A cyclothyridid rhynchonellide, a species of Burrirhynchia.
Cyclothyris tenuicostata^[58]	sp nov	Valid	Smirnova	Early Cretaceous		Russia	A cyclothyridid rhynchonellide, a species of Burrirhynchia.
Cyclothyris zudakharica^[58]	sp nov	Valid	Smirnova	Early Cretaceous		Russia	A cyclothyridid rhynchonellide, a species of Burrirhynchia.
Daghanirhynchia susanae^[55]	Species	Valid	Feldman et al.	Callovian	Mughanniyya Formation	Jordan	A tetrarhynchiid rhynchonellide, a species of Daghanirhynchia.
Duolobella^[56]	Gen. et sp. nov	Valid	Rasmussen, Harper & Blodgett	Late Ordovician	Farewell Terrane	United States	A member of Orthida belonging to the group Orthoidea and the family Orthidae. The type species is D. sandiae.
Eodmitria anauris^[61]	sp nov	Valid	Oleneva	Late Devonian (middle Frasnian)		Russia	A cyrtospiriferid spiriferid, a species of Eodmitria.
Gatosella^[62]	Gen. et sp. nov	Valid	Benedetto	Middle Ordovician (early Darriwilian)	San Juan Formation	Argentina	A plectambonitoid brachiopod. The type species is Gatosella muricata.
Gelidorthis perisiberiaensis^[56]	Sp. nov	Valid	Rasmussen, Harper & Blodgett	Late Ordovician	Farewell Terrane	United States	A member of Orthida belonging to the group Plectorthoidea and the family Plectorthidae.
Gracianella (Sublepida) paulula^[63]	sp nov	Valid	Baliński	Early Devonian	Khudykivtsi Beds	Ukraine	An atrypinid brachiopod, a species of Gracianella.
Kentronetes giae^[64]	Sp. nov	Valid	Racheboeuf in Racheboeuf et al.	Devonian (Lochkovian)	Uncía Formation	Bolivia	A chonetoid brachiopod belonging to the family Strophochonetidae.
Kuangshanotreta^[65]	Gen. et sp. nov	Valid	Wang et al.	Early Cambrian	Heilinpu Formation	China	An acrotretoid, a member of Lingulata. The type species is Kuangshanotreta malungensis.
Kudrjavzevina^[58]	Gen. et 3 sp. nov	Valid	Smirnova	Early Cretaceous		Russia	A cyclothyridid rhynchonellide. Genus contains 3 species: K. ulluchaensis, K. regularis and K. fragilis.
Lamellaerhynchia latiovalis^[58]	Sp. nov	Valid	Smirnova	Early Cretaceous		Russia	A praecyclothyridid rhynchonellide, a species of Lamellaerhynchia.
Leptaena (Septomena) alaskensis^[56]	Sp. nov	Valid	Rasmussen, Harper & Blodgett	Late Ordovician	Farewell Terrane	United States	A member of Strophomenida belonging to the family Raﬁnesquinidae.
Lobachevina^[58]	Gen. et 2 sp. nov	Valid	Smirnova	Early Cretaceous		Russia	A cyclothyridid rhynchonellide. Genus contains 2 new species: L. angusteinis and L. vagus.
Lynnica^[66]	Gen. et sp. nov	Valid	Madison	Ordovician		Russia	A member of Strophomenida. The type species is Lynnica fragilis.
Orthothrix sudoi^[67]	Sp. nov	Valid	Tazawa	Permian (Changhsingian)	Toyoma Formation	Japan	A member of Productida belonging to the suborder Strophalosiidina and the family Strophalosiidae.
Oanduporella kuskokwimensis^[56]	Sp. nov	Valid	Rasmussen, Harper & Blodgett	Late Ordovician	Farewell Terrane	United States	A member of Orthida belonging to the group Enteletoidea and the family Draboviidae.
Palaeowingella^[56]	Gen. et sp. nov	Valid	Rasmussen, Harper & Blodgett	Late Ordovician	Farewell Terrane	United States	A member of Orthida belonging to the group Orthoidea and the family Orthidae. The type species is P. farewellensis.
Plectodonta mariae pantherae^[63]	Subsp nov	Valid	Baliński	Early Devonian	Khudykivtsi Beds	Ukraine	A sowerbyellid brachiopod, a subspecies of Plectodonta mariae.
Ptychoglyptus alaensis^[56]	Sp. nov	Valid	Rasmussen, Harper & Blodgett	Late Ordovician	Farewell Terrane	United States	A member of Strophomenida belonging to the group Plectambonitoidea and the family Sowerbyellidae.
Rotutaspirifer^[61]	Gen. et sp. nov	Valid	Oleneva	Late Devonian (middle Frasnian)		Russia	A cyrtospiriferid spiriferid. The type species is Rotutaspirifer rotutus.
Sanjuanetes glemareci^[64]	Sp. nov	Valid	Racheboeuf in Racheboeuf et al.	Silurian (Přídolí)	Muruhuta Shale Member	Bolivia	A chonetoid brachiopod belonging to the family Strophochonetidae.
Sharovaella^[68]	Gen. et sp. nov	Valid	Pakhnevich	Late Devonian (Famennian)		Russia	A punctate rhynchonellide brachiopod. The type species is Sharovaella mirabilis.
Skenidioides tatyanae^[63]	sp nov	Valid	Baliński	Early Devonian	Khudykivtsi Beds	Ukraine	A skenidiid brachiopod, a species of Skenidioides.
Sowerbyella (Rugosowerbyella) praecursor^[56]	Sp. nov	Valid	Rasmussen, Harper & Blodgett	Late Ordovician	Farewell Terrane	United States	A member of Strophomenida belonging to the group Plectambonitoidea and the family Sowerbyellidae.
Sowerbyella (Sowerbyella) rectangularis^[56]	Sp. nov	Valid	Rasmussen, Harper & Blodgett	Late Ordovician	Farewell Terrane	United States	A member of Strophomenida belonging to the group Plectambonitoidea and the family Sowerbyellidae.
Strophatrypa^[69]	Gen. et sp. nov	Valid	Boucot, Blodgett & Rohr	Late Silurian		United States	A member of Atrypida belonging to the family Atrypidae. The type species is S. skaflestadi.
Sulcirhynchia insolitus^[58]	Sp. nov	Valid	Smirnova	Early Cretaceous		Russia	A praecyclothyridid rhynchonellide, a species of Sulcirhynchia.
Terrakea nabekoshiyamensis^[67]	Sp. nov	Valid	Tazawa	Permian (Changhsingian)	Toyoma Formation	Japan	A member of Productida belonging to the suborder Productidina and the family Linoproductidae.
Transridgeia^[56]	Gen. et sp. nov	Valid	Rasmussen, Harper & Blodgett	Ordovician (late Darriwilian–early Sandbian)	Farewell Terrane	United States	A member of Strophomenida belonging to the family Strophomenidae. The type species is T. costata.

Molluscs

The following is a summary of the Molluscs described in 2012

54 ammonites

14 other cephalopods

52 gastropods

15 other molluscs

Echinoderms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Anatifopsis fillmorensis^[70]	Sp. nov	Valid	Sumrall et al.	Early Ordovician		United States	A mitrate, a species of Anatifopsis.
Anatifopsis ninemilensis^[70]	Sp. nov	Valid	Sumrall et al.	Early Ordovician	McKelligon Canyon Formation Ninemile Shale	United States	A mitrate, a species of Anatifopsis.
Andymetra^[71]	Gen. et sp. nov	Valid	Hess	Early Bathonian		France	A comatulid crinoid. The type species is Andymetra galei.
Aorocrinus meyeri^[72]	Sp. nov	Valid	Ausich & Roeser	Late Kinderhookian	Cuyahoga Formation	United States	A camerate crinoid, a species of Aorocrinus.
Aphelecrinus gracilis^[73]	Sp. nov	Valid	Kammer & Roeser	Kinderhookian	Cuyahoga Formation	United States	A cladid crinoid, a species of Aphelecrinus.
Apiocrinites negevensis^[74]	Sp. nov	Valid	Ausich & Wilson	Middle Jurassic (Callovian)	Matmor Formation	Israel	An apiocrinitid crinoid, a species of Apiocrinites.
Aryballocrinus martini^[72]	Sp. nov	Valid	Ausich & Roeser	Late Kinderhookian	Cuyahoga Formation	United States	A camerate crinoid, a species of Aryballocrinus.
Atelestocrinus meszarosi^[73]	Sp. nov	Valid	Kammer & Roeser	Kinderhookian	Cuyahoga Formation	United States	A cladid crinoid, a species of Atelestocrinus.
Brissopsis (Kleinia) riccardii^[75]	Sp. nov	Valid	Parma	Eocene to Oligocene or early Miocene		Argentina	A brissopsid echinoid, a species of Brissopsis.
Calceocrinus balticensis^[76]	Sp. nov	Valid	Ausich, Wilson & Vinn	Silurian (Llandovery)		Estonia	A disparid crinoid belonging to the group Calceocrinida.
Ctenoimbricata^[77]	Gen. et sp. nov	Valid	Zamora, Rahman & Smith	Cambrian		Spain	A stem group echinoderm. The type species is Ctenoimbricata spinosa.
Cuyahogacrinus^[73]	Gen. et sp. nov	Valid	Kammer & Roeser	Kinderhookian	Cuyahoga Formation	United States	A cladid crinoid. The type species is Cuyahogacrinus lodiensis.
Cyathocrinites simplex^[73]	Sp. nov	Valid	Kammer & Roeser	Kinderhookian	Cuyahoga Formation	United States	A cladid crinoid, a species of Cyathocrinites.
Cyrtocrinus praenutans^[71]	Sp. nov	Valid	Hess	Early Bathonian		France	A cyrtocrinid crinoid, a species of Cyrtocrinus.
Desmidocrinus laevigatus^[76]	Sp. nov	Valid	Ausich, Wilson & Vinn	Silurian (Pridoli)	Kaugatuma Formation	Estonia	A camerate crinoid belonging to the group Monobathrida.
Diploblastus fadigai^[78]	Sp. nov	Valid	Atwood & Sumrall	Carboniferous (Mississippian)	Glen Dean Formation	United States	A blastoid, a species of Diploblastus.
Drepanocystis^[70]	Gen. et sp. nov	Valid	Sumrall et al.	Early Ordovician	Wah Wah Limestone	USA	A possible solute. The type species is Drepanocystis dubius.
Eucalyptocrinites tumidus^[76]	Sp. nov	Valid	Ausich, Wilson & Vinn	Silurian (Pridoli)	Kaugatuma Formation	Estonia	A camerate crinoid belonging to the group Monobathrida.
Faorina maullui^[79]	Sp. nov	Valid	Stara & Borghi	Early Miocene		Italy	A pericosmid sea urchin, a species of Faorina.
Goniocrinus sceletus^[73]	Sp. nov	Valid	Kammer & Roeser	Kinderhookian	Cuyahoga Formation	United States	A cladid crinoid, a species of Goniocrinus.
Heckerocrinus^[80]	Nom nov.	Valid	Doweld	Ordovician			A replacement name for the crinoid genus Bockia Hecker 1940.
Leadagmara^[81]	Gen. et sp. nov	Valid^[82]	Thuy et al.	Late Triassic (early Carnian)		Japan	An ophiacanthid brittle star. The type species is Leadagmara gracilispina.
Lebetocrinus ohioensis^[73]	Sp. nov	Valid	Kammer & Roeser	Kinderhookian	Cuyahoga Formation	United States	A cladid crinoid, a species of Lebetocrinus.
Neotaxocrinus^[83]	Gen. et sp. nov	Valid	Mirantsev	Late Carboniferous (Pennsylvanian)		Russia	A taxocrinid crinoid. The type species is Neotaxocrinus arendti.
Palaeocomaster messingi^[71]	Sp. nov	Valid	Hess	Early Bathonian		France	A comatulid crinoid, a species of Palaeocomaster.
Paradiabolocrinus teres^[84]	Sp. nov	Valid	Hearn & Deline	Late Ordovician	Lexington Limestone	United States	A camerate crinoid belonging to the group Diplobathra and the family Rhodocrinitidae.
Pentacrinites ausichi^[71]	Sp. nov	Valid	Hess	Late Bajocian or early Bathonian		France	A species of Pentacrinites.
Pentremites fredericki^[78]	Sp. nov	Valid	Atwood & Sumrall	Carboniferous (Mississippian)	Glen Dean Formation	United States	A blastoid, a species of Pentremites.
Pentremites meganae^[78]	Sp. nov	Valid	Atwood & Sumrall	Carboniferous (Mississippian)	Glen Dean Formation	United States	A blastoid, a species of Pentremites.
Phyllocrinus voultensis^[71]	Sp. nov	Valid	Hess	Early Bathonian		France	A cyrtocrinid crinoid, a species of Phyllocrinus.
Phymosoma ravni^[85]	Sp nov.	Valid	Schlüter et al.	Late Cretaceous		Western Europe and Middle East	A phymosomatoid sea urchin, a species of Phymosoma.
Platycrinites burkei^[72]	Sp. nov	Valid	Ausich & Roeser	Late Kinderhookian	Cuyahoga Formation	United States	A camerate crinoid, a species of Platycrinites sensu lato.
Praetetracrinus bathonicus^[71]	Sp. nov	Valid	Hess	Early Bathonian		France	A cyrtocrinid crinoid, a species of Praetetracrinus.
Pygolampas^[86]	Gen. et sp. nov	Valid	Saucède, Dudicourt & Courville	Early Cretaceous (Early Hauterivian)		France	A sea urchin. The type species is Pygolampas edita.
Saaremaacrinus^[76]	Gen. et sp. nov	Valid	Ausich, Wilson & Vinn	Silurian (Pridoli)	Kaugatuma Formation	Estonia	A camerate crinoid belonging to the group Monobathrida. The type species is S. estoniensis.
Salvaster^[86]	Gen. et sp. nov	Valid	Saucède, Dudicourt & Courville	Early Cretaceous (Early Hauterivian)		France	A sea urchin. The type species is Salvaster roberti.
Scutellacrinus^[71]	Gen. et sp. nov	Valid	Hess	Late Bajocian or early Bathonian		France	A cyrtocrinid crinoid. The type species is Scutellacrinus tenuis.
Segmentocolumnus (col.) hanshessi^[87]	Sp. nov	Valid	Donovan et al.	Middle Ordovician (latest Darriwilian)		China	Probably a stem-group cladid crinoid, possibly a dendrocrinid; a species of Segmentocolumnus.
Singillatimetra^[71]	Gen. et sp. nov	Valid	Hess	Early Bathonian		France	A comatulid crinoid. The type species is Singillatimetra inordinata.
Solanocrinites voultensis^[71]	Sp. nov	Valid	Hess	Early Bathonian		France	A comatulid crinoid, a species of Solanocrinites.
Thylechinus (Thylechinus) sinaiensis^[88]	Sp. nov	Valid	Abdelhamid & Azab	Turonian		Egypt	A phymosomatid echinoid, a species of Thylechinus.
Toxaster dakhlensis^[88]	Sp. nov	Valid	Abdelhamid & Azab	Turonian		Egypt	A toxasterid echinoid, a species of Thylechinus.
Tripatagus^[89]	Gen. et sp. nov	Valid	Zachos	Oligocene (Rupelian)	Marianna Limestone	United States	A heart urchin belonging to the group Micrasterina. The type species is T. pittsi.
Yanjiahella^[90]	Gen. et 3 sp. nov	Valid	Guo et al.	Early Cambrian	Yanjiahe Formation	China	An animal of uncertain phylogenetic position, a possible relative of echinoderms. Genus contains Yanjiahella ancarpa (the type species), Y. monocarpa and Y. biscarpa.

Ascidians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Burykhia^[91]	Gen. et sp.	Valid	Fedonkin et al.	Vendian	Verkhovka Formation or Ust-Pinega Formation	Russia	An ausiid, a possible ascidian, Its type species is Burykhia hunti.

Conodonts

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Carnepigondolella angulata^[92]	Sp nov	Valid	Mazza, Cau & Rigo	Late Triassic		Canada^[93] Italy	Originally described as a species of Carnepigondolella; Orchard (2014) transferred this species to the genus Quadralella.^[93]
Carnepigondolella gulloae^[94]	Sp nov	Valid	Mazza, Rigo & Gullo	Late Triassic	Scillato Formation	Italy	A species of Carnepigondolella.
Carnepigondolella tuvalica^[94]	Sp nov	Valid	Mazza, Rigo & Gullo	Late Triassic	Scillato Formation	Italy	A species of Carnepigondolella.
Caudicriodus schoenlaubi^[95]	Sp nov	Valid	Drygant & Szaniawski	Early Devonian (middle to late Lochkovian)	Chortkiv Formation	Ukraine	An icriodontid, a species of Caudicriodus.
Epigondolella heinzi^[92]	Sp nov	Valid	Mazza, Cau & Rigo	Late Triassic		Italy	A species of Epigondolella.
Epigondolella miettoi^[92]	Sp nov	Valid	Mazza, Cau & Rigo	Late Triassic		Italy	A species of Epigondolella.
Idiognathodus espinamaenis^[96]	Sp. nov	Valid	Méndez	Carboniferous (Moscovian)		Spain
Idiognathodus mendezi^[96]	Sp. nov	Valid	Méndez	Carboniferous (Moscovian)		Spain
Meiognathus^[97]	Gen. et sp. nov	Valid	Shen et al.	Permian (Kungurian)		Japan	A member of Ozarkodinida belonging to the family Sweetognathidae. The type species is Meiognathus pustulus.
Muellerina^[98]	Gen. et sp. nov	Junior homonym	Bardashev & Bardasheva	Middle Devonian (Givetian)		Tajikistan	The type species is Muellerina idrisovi. The generic name turned out to be preoccupied by Muellerina Bassiouni (1965); Bardashev & Bardasheva (2013) subsequently renamed the conodont genus Muellerilepis.^[99]
Nealeodus^[100]	Gen. et comb. nov	Valid	Stouge	Middle Ordovician		Canada	A new genus for "Lenodus" martinpointensis (Johnston and Barnes, 2000).
Norigondolella trinacriae^[92]	Sp nov	Valid	Mazza, Cau & Rigo	Late Triassic		Italy	Originally described as a species of Norigondolella; Karádi, Kozur & Görögt (2013) considered it more likely to be a species of Paragondolella.^[101]
Pandorinellina? parva^[95]	Sp nov	Valid	Drygant & Szaniawski	Early Devonian (late Lochkovian or Pragian)	Ivanye Formation	Ukraine	A spathognathodontid, possibly a species of Pandorinellina.
Periodon hankensis^[100]	Sp nov	Valid	Stouge	Middle Ordovician		Canada	A species of Periodon.
Polygnathus chongqingensis^[102]	Sp. nov	Valid	Wang in Gong et al.	Devonian		China
Pseudopolygnathus inordinatus^[103]	Nom. nov	Valid	Tragelehn & Hartenfels	Devonian (Famennian)		Germany	A replacement name for Pseudopolygnathus irregularis Tragelehn & Hartenfels (2011) (preoccupied by Pseudopolygnathus irregularis Branson, 1934).
Spinodus wardi^[100]	Sp nov	Valid	Stouge	Middle Ordovician		Canada	A species of Spinodus.

Fishes

During 2012, 81 new species of fish were described.

Amphibians

Research

A study of the braincase of Eocaecilia micropodia and a phylogenetic analysis of non-amniote tetrapods is published by Hillary C. Maddin, Farish A. Jenkins Jr and Jason S. Anderson (2012).^[104]

A study of anatomy and relationships of Solenodonsaurus janenschi is published by Marylène Danto, Florian Witzmann and Johannes Müller (2012).^[105]

A study of limb joint mobility of Ichthyostega is published by Stephanie E. Pierce, Jennifer A. Clack and John R. Hutchinson (2012).^[106]

New taxa

Newly named basal tetrapods

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Ymeria^[107]	Gen. et sp. nov	Valid	Clack, Ahlberg, Blom & Finney	Famennian	Celsius Bjerg Group	Greenland	A stem tetrapod closely related to Ichthyostega.

Newly named temnospondyls

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Arachana^[108]	Gen. et sp. nov	Valid	Piñeiro, Ramos & Marsicano	? Late Permian	Buena Vista Formation	Uruguay	A rhinesuchid-like stereospondyl.
Benthosuchus gusevae^[109]	Sp. nov	Valid	Novikov	Early Triassic	Kamennyi Yar Formation	Russia	A trematosauroid.
Cacops woehri^[110]	Sp. nov	Valid	Fröbisch & Reisz	Leonardian		USA	A dissorophid.
Microposaurus averyi^[111]	Sp nov	Valid	Warren	Anisian	Rouse Hill Siltstone	Australia	A trematosaurine stereospondyl, a species of Microposaurus.	Microposaurus averyi.
Nyranerpeton^[112]	Gen. et sp. nov	Valid	Werneburg	Late Carboniferous		Czech Republic	A micromelerpetontid.^[113] The type species is Nyranerpeton amilneri.^[112]
Platyrhinops fritschi^[112]	Sp. nov	Valid	Werneburg	Late Carboniferous		Czech Republic	An amphibamid, a species of Platyrhinops.
Qantas^[114]	Gen. et sp. nov	Valid	Novikov	Early Triassic	Kamennyi Yar Formation	Russia	A trematosauroid temnospondyl. The type species is Qantas samarensis.
Sclerocephalus stambergi^[115]	Sp nov	Valid	Klembara & Steyer	Early Permian	Boskovice Basin	Czech Republic	A stereospondylomorph, a species of Sclerocephalus.
Trematosuchoides^[114]	Gen. et sp. nov	Valid	Novikov	Triassic	Cynognathus Assemblage Zone	South Africa	A trematosaurid temnospondyl. The type species is Trematosuchoides africanus.

Newly named lepospondyls

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Altenglanerpeton^[116]	Gen. et sp. nov	Valid	Glienke	Gzhelian or Asselian	Altenglan Formation	Germany	A microsaur with an elongated body.

Newly named lissamphibians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Albionbatrachus oligocenicus^[117]	Sp. nov	Valid^[118]	Venczel, Codrea & Fărcaş	Early Oligocene		Romania	A palaeobatrachid frog, a species of Albionbatrachus.
Bakonybatrachus^[119]	Gen. et sp. nov	Valid	Szentesi & Venczel	Santonian	Csehbánya Formation	Hungary	A discoglossine discoglossid.
Beiyanerpeton^[120]	Gen. et sp. nov	Valid	Gao & Shubin	Oxfordian	Tiaojishan Formation	China	A salamandroid. The type species is Beiyanerpeton jianpingensis.
Calyptocephalella satan^[121]	Sp. nov	Valid	Agnolin	Late Cretaceous (Campanian-Maastrichtian)	Allen Formation	Argentina	A member of Calyptocephalellidae, a relative of the helmeted water toad.
Gigantobatrachus casamiquelai^[121]	Sp. nov	Valid	Agnolin	Paleocene		Argentina	A member of Calyptocephalellidae, a species of Gigantobatrachus.
Piceoerpeton naylori^[122]	Sp. nov.	Valid	Gardner	Latest Cretaceous (late Maastrichtian), possibly also early Paleocene		United States	A scapherpetontid salamander, a species of Piceoerpeton.
Rana auscitana^[123]	Nom. nov	Valid	Martín, Alonso-Zarazaga & Sanchiz	Middle Miocene		France	A replacement name for Rana pygmaea Lartet (1851).
Rana cadurcorum^[123]	Nom. nov	Junior synonym	Martín, Alonso-Zarazaga & Sanchiz	Eocene	Quercy Phosphorites	France	A replacement name for Rana plicata Filhol (1877). Reinterpreted as a junior synonym of Thaumastosaurus gezei by Laloy et al. (2013).^[124]
Rana sendoa^[123]	Nom. nov	Valid	Martín, Alonso-Zarazaga & Sanchiz	Late Pleistocene		Germany	A replacement name for Rana robusta Brunner (1956).
Seminobatrachus^[125]	Gen. et sp. nov	Valid	Skutschas & Gubin	late Paleocene - early Eocene		Ukraine	A urodelan salamander
Uberabatrachus^[126]	Gen. et sp. nov	Valid	Báez et al.	Maastrichtian	Marília Formation	Brazil	A neobatrachian, a possible member of Nobleobatrachia. The type species is Uberabatrachus carvalhoi.

Parareptiles

Newly named parareptiles

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Belebey augustodunensis^[127]	Sp. nov.	Valid	Falconnet	Latest Carboniferous (Gzhelian) or earliest Permian (Asselian)		France	A bolosaurid, a species of Belebey.
Feeserpeton^[128]	Gen. et sp. nov.	Valid	Macdougall & Reisz	Early Permian		United States	A lanthanosuchoid. The type species is Feeserpeton oklahomensis.
Lasasaurus^[129]	Gen. et sp. nov.	Valid	Falconnet et al.	Lower Triassic	Middle Sakamena Formation	Madagascar	A procolophonid. The type species is Lasasaurus beltanae.

Ichthyopterygians

Newly named ichthyosaurs

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Acamptonectes^[130]	Gen. et sp. nov	Valid	Fischer et al..	Hauterivian–Cenomanian	Cambridge Greensand Formation	United Kingdom	An ophthalmosaurid. Type species is Acamptonectes densus.	Skull roof of Acamptonectes densus.
Cryopterygius^[131]	Gen. et sp. nov	Valid	Druckenmiller et al.	Late Jurassic	Agardhfjellet Formation	Norway	An ophthalmosaurid. The type species is Cryopterygius kristiansenae.
Palvennia^[131]	Gen. et sp. nov	Valid	Druckenmiller et al.	Late Jurassic	Agardhfjellet Formation	Norway	An ophthalmosaurid. The type species is Palvennia hoybergeti.
Stenopterygius aaleniensis^[132]	Sp. nov	Valid	Maxwell, Fernández & Schoch	Aalenian		Germany	A species of Stenopterygius.	The holotype of Stenopterygius aaleniensis.
Temnodontosaurus azerguensis^[133]	Sp. nov	Valid	Martin et al.	Middle Toarcian		France	A species of Temnodontosaurus.

Lepidosauromorphs

Newly named saurosphargids

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Largocephalosaurus^[134]	Gen. et sp. nov	Valid	Cheng, Chen, Zeng & Cai	Middle Triassic	Guanling Formation	China	Initially thought to be an eosauropterygian sauropterygian and a relative of Wumengosaurus, pachypleurosaurs and nothosauroids,^[134] but subsequently reinterpreted as member of Saurosphargidae (a non-sauropterygian diapsid, though related to sauropterygians), closely related to Saurosphargis and Sinosaurosphargis.^[135] The type species is Largocephalosaurus polycarpon.

Newly named sauropterygians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Albertonectes^[136]	Gen. et sp. nov	Valid	Kubo, Mitchell & Henderson	Upper Campanian	Bearpaw Formation	Canada	An elasmosaurid. The type species is Albertonectes vanderveldei.	Albertonectes vanderveldei.
Anningasaura^[137]	Gen. et sp. nov.	Valid	Vincent & Benson	Early Jurassic		United Kingdom	A basal member of Plesiosauria. The type species is Anningasaura lymense.
Avalonnectes^[138]	Gen. et sp. nov	Valid	Benson, Evans & Druckenmiller	Early Jurassic, most likely earliest Hettangian.	Blue Lias Formation	United Kingdom	A rhomaleosaurid. The type species is Avalonnectes arturi.	Avalonnectes.
Djupedalia^[139]	Gen. et sp. nov.	Valid	Knutsen, Druckenmiller & Hurum	Late Jurassic	Agardhfjellet Formation	Norway	A long-necked plesiosaurian. The type species is Djupedalia engeri.
Dolichorhynchops tropicensis^[140]	sp nov	Valid	McKean	Early Turonian	Tropic Shale	United States	Third species of Dolichorhynchops.
Eoplesiosaurus^[138]	Gen. et sp. nov	Valid	Benson, Evans & Druckenmiller	Early Jurassic, most likely earliest Hettangian.		United Kingdom	A basal plesiosauroid. The type species is Eoplesiosaurus antiquior.	Eoplesiosaurus.
Hastanectes^[141]	Gen. et comb. nov	Valid^[142]	Benson et al.	Early Cretaceous		United Kingdom	A new genus for "Cimoliasaurus" valdensis
Lusonectes^[143]	Gen. et sp. nov	Valid	Smith, Araújo & Mateus	Toarcian	São Gião Formation	Portugal	A plesiosaurid plesiosaur.
Pliosaurus funkei^[144]	Sp. nov	Valid	Knutsen, Druckenmiller & Hurum	Late Jurassic	Agardhfjellet Formation	Norway	A species of Pliosaurus.
Qianxisaurus^[145]	Gen. et sp. nov	Valid	Cheng et al.	Middle Triassic (Ladinian)	Falang Formation	China	An eosauropterygian, a relative of pachypleurosaurs and nothosauroids. The type species is Qianxisaurus chajiangensis.
Spitrasaurus^[146]	Gen. et 2 sp. nov.	Valid	Knutsen, Druckenmiller & Hurum	Late Jurassic	Agardhfjellet Formation	Norway	A long-necked plesiosaurian. Genus contains two species: Spitrasaurus wensaasi and S. larseni.
Stratesaurus^[138]	Gen. et sp.	Valid	Benson, Evans & Druckenmiller	Early Jurassic, most likely earliest Hettangian.	Blue Lias Formation	United Kingdom	A rhomaleosaurid. The type species is Stratesaurus taylori	Stratesaurus.
Vectocleidus^[141]	Gen. et sp. nov	Valid^[142]	Benson et al.	Early Cretaceous (late Barremian)	Vectis Formation	United Kingdom	A leptocleidid. The type species is Vectocleidus pastorum.

Newly named rhynchocephalians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Oenosaurus^[147]	Gen. et sp. nov	Valid	Rauhut et al.	Late Jurassic (early Tithonian)	Mörnsheim Formation	Germany	A sphenodontid rhynchocephalian. The type species is Oenosaurus muehlheimensis.
Sphenocondor^[148]	Gen. et sp. nov	Valid	Apesteguía, Gómez & Rougier	Middle Jurassic		Argentina	A rhynchocephalian lepidosaur. The type species is Sphenocondor gracilis.

Squamates

Research

A large phylogenetic analysis of living and fossil squamates is published by Jacques A. Gauthier et al. (2012).^[149]

A study of squamate diversity in North America during the latest Cretaceous (Maastrichtian) and the impact of Cretaceous–Paleogene extinction event on the diversity of the group is published by Nicholas R. Longrich, Bhart-Anjan S. Bhullar and Jacques A. Gauthier (2012).^[150]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Chianghsia^[151]	Gen. et sp. nov	Valid	Mo, Xu & Evans	Late Cretaceous	Nanxiong Formation	China	A platynotan lizard, probably a monstersaurian. The type species is Chianghsia nankangensis.
Dornosaurus^[152]	Gen. et sp. nov	Valid	Alifanov	Eocene		Mongolia	An arretosaurid iguanian. The type species is Dornosaurus gobiensis.
Eremiasaurus^[153]	Gen. et sp. nov	Valid	Leblanc, Caldwell & Bardet	Maastrichtian		Morocco	A mosasaur.
Ergiliinsaurus^[152]	Gen. et sp. nov	Valid	Alifanov	Oligocene	Ergiliin Zoo Formation	Mongolia	An arretosaurid iguanian. The type species is Ergiliinsaurus postumus.
Jucaraseps^[154]	Gen. et sp. nov	Valid	Bolet & Evans	Early Cretaceous		Spain	A lizard related to scleroglossan lizards. The type species is Jucaraseps grandipes.
Khaichinguana^[152]	Gen. et sp. nov	Valid	Alifanov	Eocene	Khaychin Formation	Mongolia	An arretosaurid iguanian. The type species is Khaichinguana eocaenica.
Louisamphisbaena^[155]	Gen. et sp. nov	Valid	Augé	Eocene (Bartonian)		France	An amphisbaenian lizard, possibly a member of Blanidae. The type species is Louisamphisbaena ferox.
Pannoniasaurus^[156]	Gen. et sp. nov	Valid	Makádi, Caldwell & Ősi	Late Cretaceous (Santonian)	Ajka Coal Formation Csehbánya Formation	Hungary	A tethysaurine mosasauroid closely related to Tethysaurus, Russellosaurus and Yaguarasaurus. The type species is Pannoniasaurus inexpectatus.
?Placosaurus ragei^[157]	Sp. nov.	Valid	Sullivan et al.	Earliest Eocene		Belgium	A glyptosaurine anguid lizard, possibly a species of Placosaurus.
Pseudomimeosaurus^[158]	Gen. et comb. nov	Valid	Alifanov	Late Cretaceous	Djadochta Formation	Mongolia	A pleurodontagamid iguanian lizard; a new genus for "Mimeosaurus" tugrikinensis Alifanov (1989).
Pseudopus ahnikoviensis^[159]	Sp. nov	Valid	Klembara	Early Miocene		Czech Republic	An anguid, a species of Pseudopus.
Purbicella^[160]	Gen. et sp. nov.	Valid	Evans, Jones & Matsumoto	Early Cretaceous	Purbeck Limestone Group	United Kingdom	A member of Lacertoidea. The type species isPurbicella ragei.
Ragesaurus^[161]	Gen. et sp. nov.	Valid	Bailon & Auge	Early Pleistocene		Spain	An anguid lizard. The type species is Ragesaurus medasensis.
Sphaerodactylus ciguapa^[162]	Sp. nov	Valid	Daza & Bauer	Late Early Miocene or early Middle Miocene (20 to 15 MYA)		Dominican Republic	A sphaerodactylid gecko found in Dominican amber, a species of Sphaerodactylus.
Telmasaurus bialynickae^[158]	Sp. nov	Valid	Alifanov	Late Cretaceous	Barun Goyot Formation	Mongolia	A varanoid lizard, a species of Telmasaurus.
Uquiasaurus^[163]	Gen. et sp. nov	Valid	Daza et al.	Late Pliocene	Uquía Formation	Argentina	An iguanian lizard. The type species is Uquiasaurus heptanodonta.
Varanus (Varaneades) amnhophilis^[164]	Subgen. et sp. nov	Disputed	Conrad, Balcarcel & Mehling	Miocene (Turolian)	Mytilini Formation	Greece	A monitor lizard. Villa et al. (2018) considered this species to be likely junior synonym of Varanus marathonensis.^[165]

Turtles

Research

A large phylogenetic analysis of basal turtles is published by Jérémy Anquetin (2012).^[166]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Allopleuron qazaqstanense^[167]	Sp. nov	Valid	Karl, Gröning & Brauckmann	Eocene (Lutetian)		Kazakhstan	A member of Cheloniidae, a species of Allopleuron.
Axestemys cerevisia^[168]	sp nov	Valid	Vitek	Eocene	Bridger Formation	United States	A trionychid, a species of Axestemys.
Axestemys montinsana^[168]	sp nov	Valid	Vitek	Paleocene	Denver Formation Fort Union Formation Melville Formation	United States	A trionychid, a species of Axestemys.
Ballerstedtia^[169]	Gen. et comb. et sp. nov	Valid	Karl et al.	Early Cretaceous		Germany United Kingdom	A member of Paracryptodira, a new genus for "Pleurosternon" typocardium Seeley (1869). Genus also contains a new species Ballerstedtia bueckebergensis.
Bashuchelys^[170]	Gen. et comb. et sp. nov	Valid	Tong, Danilov, Ye, Ouyang & Peng	Middle Jurassic	Xiashaximiao Formation	China	A bashuchelyid cryptodiran, a new genus for "Chengyuchelys" zigongensis (Ye, 1982). Genus also contains a new species Bashuchelys youngi.
Berruchelus^[171]	Gen. et sp. nov	Valid	Pérez-García	Upper Paleocene		France	A paracryptodiran closely related to Compsemys. The type species is Berruchelus russelli.
Carbonemys^[172]	Gen. et sp. nov	Valid	Cadena et al.	Late Palaeocene	Cerrejón Formation	Colombia	A podocnemidid turtle. The type species is Carbonemys cofrinii.
Chuannanchelys^[170]	Gen. et comb. nov	Valid	Tong, Danilov, Ye, Ouyang & Peng	Middle Jurassic	Xiashaximiao Formation	China	A bashuchelyid cryptodiran, a new genus for "Chengyuchelys" dashanpuensis (Fang, 1987).
Foxemys trabanti^[173]	sp nov	Valid	Rabi, Tong & Botfalvai	Santonian	Csehbánya Formation	Hungary	A bothremydid, a species of Foxemys.
Galvechelone^[174]	Gen. et sp. nov	Valid	Pérez-García & Murelaga	Early Cretaceous		Spain	A cryptodiran turtle. The type species is Galvechelone lopezmartinezae.
Guangdongemys^[175]	Gen. et sp. nov.	Valid	Claude et al.	Late Eocene		China	A geoemydid turtle. The type species is Guangdongemys pingi.
Hoyasemys^[176]	Gen. et sp. nov	Valid	Pérez-García, Fuente & Ortega	Late Barremian	Calizas de La Huérguina Formation	Spain	A basal eucryptodiran.
Iberoccitanemys^[177]	Gen. et comb. nov	Valid	Pérez-García, Ortega & Murelaga	Late Campanian to Maastrichtian		France Spain	A bothremydid, a new genus for "Elochelys" convenarum (Laurent et al. 2002).
Kinosternon pojoaque^[178]	Species	Valid	Bourque	Middle Miocene		United States	A species of Kinosternon.
Kinosternon skullridgescens^[179]	Species	Valid	Bourque	Early middle Miocene	Tesuque Formation	United States	A species of Kinosternon.
Larachelus^[180]	Gen. et sp. nov	Valid	Pérez-García & Murelaga	Early Cretaceous (late Hauterivian–early Barremian)	Pinilla de los Moros Formation	Spain	A basal member of Pan-Cryptodira (the clade containing living cryptodirans and all extinct turtles that were more closely related to them than to pleurodirans). The type species is Larachelus morla.
Oertelia^[181]	Gen. et comb. nov	Valid	Karl, Biermann & Tichy	Early Cretaceous (early Aptian)		Germany	A toxochelyid sea turtle, a new genus for "Toxochelys" gigantea Oertel (1914).
Peligrochelys^[182]	Gen. et sp. nov	Valid^[183]	Sterli & de la Fuente	Palaeocene	Salamanca Formation	Argentina	A relative of meiolaniids. The type species is Peligrochelys walshae.
Polysternon isonae^[184]	Species	Valid	Marmi et al..	Late Maastrichtian		Spain	A bothremydid, a species of Polysternon.
Protoxinjiangchelys^[170]	Gen. et sp. nov	Valid	Tong, Danilov, Ye, Ouyang & Peng	Middle Jurassic	Xiashaximiao Formation	China	A xinjiangchelyid cryptodiran.
Puentemys^[185]	Gen. et sp. nov.	Valid	Cadena, Bloch & Jaramillo	Paleocene	Cerrejón Formation	Colombia	A bothremydid turtle. The type species is Puentemys mushaisaensis.
Rhinoclemmys panamaensis^[186]	Sp nov	Valid	Cadena et al.	Miocene	Cucaracha Formation	Panama	A geoemydid turtle, a species of Rhinoclemmys.
Staurotypus moschus^[186]	Sp nov	Valid	Cadena et al.	Miocene	Cucaracha Formation	Panama	A kinosternid turtle, a species of Staurotypus.
Tasbacka danica^[187]	Sp. nov	Valid	Karl & Madsen	Early Eocene	Fur Formation	Denmark	A member of Cheloniidae, a species of Tasbacka.
Terrapene parornata^[188]	Sp nov	Valid	Joyce et al..	Miocene/Pliocene boundary		United States	A box turtle.
Yuchelys^[189]	Gen. et sp. nov	Valid	Tong et al.	Late Cretaceous	Gaogou Formation	China	A nanhsiungchelyid cryptodiran. The type species is Yuchelys nanyangensis.

Archosauromorphs

Newly named basal archosauromorphs

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Chanaresuchus ischigualastensis^[190]	sp nov	Valid	Trotteyn, Martínez & Alcober	Late Triassic	Ischigualasto Formation	Argentina	A proterochampsian archosauriform. Originally described as a species of Chanaresuchus;^[190] subsequently made the type species of a separate genus Pseudochampsa by Trotteyn & Ezcurra (2014).^[191]
Diandongosuchus^[192]	Gen. et sp. nov.	Valid	Li et al.	Middle Triassic (Ladinian)	Falang Formation	China	A member of Archosauriformes of uncertain phylogenetic placement. Originally classified as a poposauroid pseudosuchian,^[192] but subsequently argued to be a phytosaur.^[193]^[194] The type species is Diandongosuchus fuyuanensis.	Diandongosuchus fuyuanensis.
Doswellia sixmilensis^[195]	sp nov	Valid	Heckert, Lucas & Spielmann	Late Triassic	Bluewater Creek Formation	United States	A doswelliid archosauromorph, a species of Doswellia.
Protome^[196]	Gen. et sp. nov	Valid	Stocker	Late Triassic	Chinle Formation	USA	A phytosaur. The type species is Protome batalaria.

Archosaurs

The following is a summary of the archosaurs described in 2012

8 pseudosuchians

1 basal dinosauriform

40 non-avian dinosaurs

64 birds

9 pterosaurs

1 archosaur of uncertain phylogenetic placement

Synapsids

Non-mammalian synapsids

Research

A phylogenetic analysis of basal synapsids is published by Roger B. J. Benson (2012).^[197]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Fortunodon^[198]	Gen. et comb. nov	Valid	Kurkin	Upper Permian		Russia	A dicynodont, a new genus for "Dicynodon" trautscholdi (Amalitzky, 1922) (considered to be a senior synonym of Vivaxosaurus permirus and assigned to the genus Vivaxosaurus by Kammerer et al., 2011).^[199]"Dicynodon" amalitzkii (Sushkin, 1926) is considered by Kurkin (2012) to be a second species of Fortunodon^[198] (Kammerer et al., 2011 classify it as the type species of the genus Peramodon.^[199])
Novocynodon^[200]	Gen. et sp. nov	Valid	Ivakhnenko	Middle Permian		Russia	A thrinaxodontid cynodont. The type species is Novocynodon kutorgai.
Pampaphoneus^[201]	Gen. et sp. nov	Valid	Cisneros, Abdala, et al.	Guadalupian (Middle Permian)	Rio do Rasto Formation	Brazil	An anteosaurid dinocephalian.
Panchetocynodon^[202]	Gen. et sp. nov	Valid	Das & Gupta	Early Triassic	Panchet Formation	India	A cynodont. The type species is Panchetocynodon damodarensis.
Purbeckodon^[203]	Gen. et sp. nov	Valid	Butler, Sigogneau-Russell, & Ensom	Early Cretaceous	Purbeck Limestone Group	United Kingdom	A possible morganucodontan.
Sludica^[200]	Gen. et sp. nov	Valid	Ivakhnenko	Upper Permian		Russia	A procynosuchid cynodont. The type species is Sludica bulanovi.

Mammals

The following is a summary of the mammals described in 2012

9 non-eutherian Mammals

24 eutherian Mammals

Other animals

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Aggerscolex^[204]	Gen. et sp. nov	Valid	Botting et al.	Ordovician		United Kingdom	A palaeoscolecid. The type species is A. murchisoni.
Bullascolex^[204]	Gen. et sp. nov	Valid	Botting et al.	Ordovician		United Kingdom	A palaeoscolecid. The type species is B. inserere.
Carbotubulus^[205]	Gen. et sp. nov	Valid	Haug et al.	Carboniferous	Carbondale Formation	United States	A long-legged lobopodian. The type species is C. waloszeki.
Circotheca smetanai^[206]	Sp. nov	Valid	Valent et al.	Middle Cambrian	Buchava Formation	Czech Republic	A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Circotheca.
Coronacollina^[207]	Gen. et sp. nov.	Valid	Clites, Droser & Gehling	Ediacaran		Australia	An animal of uncertain phylogenetic placement. The type species is Coronacollina acula.
Crenulipora kuwanoi^[208]	Sp. nov	Valid	Niko	Devonian	Fukuji Formation	Japan	A tabulate coral belonging to the group Favositida and the family Favositidae.
Dispinoscolex^[209]	Gen. et sp. nov	Valid	Duan, Dong & Donoghue	Cambrian (Paibian)	Bitiao Formation	China	A palaeoscolecid. The type species is D. decorus.
Favia maitreyiae^[210]	Sp. nov	Valid	López-Pérez	Pliocene	Carmen Formation	Mexico	A stony coral, a species of Favia.
Favia tulsidasi^[210]	Sp. nov	Valid	López-Pérez	Pliocene	San Marcos Formation	Mexico	A stony coral, a species of Favia.
Heydenius arachnius^[211]	Sp. nov	Valid	Poinar	Eocene to Miocene	Dominican amber	Dominican Republic	A nematode belonging to the family Mermithidae.
Heydenius phasmatophilus^[211]	Sp. nov	Valid	Poinar	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A nematode belonging to the family Mermithidae.
Heydenius podenasae^[211]	Sp. nov	Valid	Poinar	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A nematode belonging to the family Mermithidae.
Heydenius trichorosus^[211]	Sp. nov	Valid	Poinar	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A nematode belonging to the family Mermithidae.
Loriciscolex^[204]	Gen. et sp. nov	Valid	Botting et al.	Ordovician		United Kingdom	A palaeoscolecid. The type species is L. cuspidus.
Parastriatopora boliviana^[64]	Sp. nov	Valid	Plusquellec, Tourneur & Fernández-Martínez in Racheboeuf et al.	Devonian (Emsian or Eifelian)	Belén Formation	Bolivia	A tabulate coral belonging to the superfamily Pachyporoidea and the family Parastriatoporidae.
Placosmilia? aliciae^[210]	Sp. nov	Valid	López-Pérez	Pliocene	San Marcos Formation San Nicolas Formation	Mexico	A stony coral belonging to the family Montlivaltiidae.
Pluoscolex^[204]	Gen. et sp. nov	Valid	Botting et al.	Ordovician		United Kingdom	A palaeoscolecid. The type species is P. linearis.
Probactrotheca^[206]	Gen. et sp. nov	Valid	Valent et al.	Middle Cambrian	Buchava Formation	Czech Republic	A member of Hyolitha. The type species is Probactrotheca briketa.
Protomichelinia foveata^[212]	Sp. nov	Valid	Niko, Haikawa & Fujikawa	Permian	Akiyoshi Limestone Group	Japan	A tabulate coral.
Pseudoromingeria muratai^[212]	Sp. nov	Valid	Niko, Haikawa & Fujikawa	Permian	Akiyoshi Limestone Group	Japan	A tabulate coral.
Radnorscolex^[204]	Gen. et sp. nov	Valid	Botting et al.	Ordovician		United Kingdom	A palaeoscolecid. The type species is R. bwlchi.
Redondagnathus^[213]	Gen. et sp. nov	Valid	Spielmann & Lucas	Late Triassic		United States	A vertebrate of uncertain phylogenetic placement. Described on the basis of three postcanine teeth, considered to be teeth of a trirachodontid cynodont by Lucas et al. (1999)^[214] and Spielmann and Lucas (2012).^[213] On the other hand, Abdala, Neveling and Welman (2006) and Sidor and Hopson (2018) did not consider it likely that Redondagnathus was a member of Trirachodontidae, and Abdala, Neveling and Welman (2006) weren't even sure whether it was a cynodont.^[215]^[216] The type species is Redondagnathus hunti.
Schistoscolex hunanensis^[209]	Sp. nov	Valid	Duan, Dong & Donoghue	Cambrian (Paibian)	Bitiao Formation	China	A palaeoscolecid.
Siderastrea annae^[210]	Sp. nov	Valid	López-Pérez	Pliocene	San Marcos Formation	Mexico	A stony coral, a species of Siderastrea.
Sinopathes^[217]	Gen. et sp. nov	Valid	Baliński, Sun & Dzik	Early Ordovician	Fenxiang Formation	China	A black coral. The type species is S. reptans.
Siphusauctum^[218]	Gen. et sp. nov	Valid	O'Brien & Caron	Middle Cambrian	Burgess Shale	Canada	A stem-bilaterian of uncertain phylogenetic placement, a stalked filter feeder. The type species is S. gregarium.
Syringopora konishii^[219]	Sp. nov	Valid	Niko & Ibaraki	Carboniferous (Viséan)	Kotaki Formation	Japan	A tabulate coral belonging to the group Auloporida.
Thecia ichikawai^[220]	Sp. nov	Valid	Niko & Adachi	Silurian	Gionyama Formation	Japan	A tabulate coral belonging to the group Favositida and the family Theciidae.
Trachypsammia konbo^[221]	Sp. nov	Valid	Niko	Early Carboniferous	Akiyoshi Limestone Group	Japan	A tabulate coral.
Ulexiscolex^[204]	Gen. et sp. nov	Valid	Botting et al.	Ordovician		United Kingdom	A palaeoscolecid. The type species is U. ormrodi.
Wernia^[204]	Gen. et sp. nov	Valid	Botting et al.	Ordovician		United Kingdom	A palaeoscolecid. The type species is W. eximia.

Other organisms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Hydrolithon braganum^[222]	Sp. nov	Valid	Woelkerling, Bassi & Iryu	Quaternary		Reef terraces around Tahiti, French Polynesia	A red alga belonging to the family Corallinaceae, a species of Hydrolithon.
Jimwhitea^[223]	Gen. et sp. nov	Valid	Krings & Taylor in Krings et al.	Middle Triassic	Fremouw Formation	Antarctica	A fungus belonging to the group Endogonales. The type species is Jimwhitea circumtecta.

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^ Juan L. Benedetto (2012). "Gatosella, a new basal plectambonitoid brachiopod with undercut cardinal process from Middle Ordovician limestones of the Precordillera terrane, Argentina". Journal of Systematic Palaeontology. 10 (3): 435–443. doi:10.1080/14772019.2011.590536.

^ ^a^b^c Andrzej Baliński (2012). "The brachiopod succession through the Silurian–Devonian boundary beds at Dnistrove, Podolia, Ukraine". Acta Palaeontologica Polonica. 57 (4): 897–924. doi:10.4202/app.2011.0138.

^ ^a^b^c Patrick R. Racheboeuf; Jean-Georges Casier; Yves Plusquellec; Margarita Toro; David Mendoza; Maria da Gloria Pires de Carvalho; Alain Le Hérissé; Florentin Paris; Esperanza Fernández-Martinez; Francis Tourneur; Jean Broutin; Sylvie Crasquin; Philippe Janvier (2012). "New data on the Silurian–Devonian palaeontology and biostratigraphy of Bolivia". Bulletin of Geosciences. 87 (2): 269–314. doi:10.3140/bull.geosci.1248.

^ Haizhou Wang; Zhifei Zhang; Lars E. Holmer; Shixue Hu; Xiangren Wang; Guoxiang Li (2012). "Peduncular attached secondary tiering acrotretoid brachiopods from the Chengjiang fauna: Implications for the ecological expansion of brachiopods during the Cambrian explosion". Palaeogeography, Palaeoclimatology, Palaeoecology. 323–325: 60–67. doi:10.1016/j.palaeo.2012.01.027.

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^ ^a^b^c Colin D. Sumrall; James Sprinkle; Thomas E. Guensburg; Benjamin F. Dattilo (2012). "Early Ordovician Mitrates and A Possible Solute (Echinodermata) from the Western United States". Journal of Paleontology. 86 (4): 595–604. doi:10.1666/10-165R.1.

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[69] Arthur J. Boucot; Robert B. Blodgett; David M. Rohr (2012). "Strophatrypa, a new genus of Brachiopoda (Atrypidae), from upper Silurian strata of the Alexander terrane, northeast Chichagof Island, Alaska". Bulletin of Geosciences. 87 (2): 261–267. doi:10.3140/bull.geosci.1285.

[JPSumralletal-70] Colin D. Sumrall; James Sprinkle; Thomas E. Guensburg; Benjamin F. Dattilo (2012). "Early Ordovician Mitrates and A Possible Solute (Echinodermata) from the Western United States". Journal of Paleontology. 86 (4): 595–604. doi:10.1666/10-165R.1.

[SJPHess-71] ⁱ Hans Hess (2012). "Crinoids from the Middle Jurassic (Bajocian–Lower Callovian) of Ardèche, France". Swiss Journal of Palaeontology. 131 (2): 211–253. doi:10.1007/s13358-012-0044-9.

[CamerateandDisparidCrinoidsJP-72] William I. Ausich; Edgar W. Roeser (2012). "Camerate and Disparid Crinoids from the Late Kinderhookian Meadville Shale, Cuyahoga Formation of Ohio". Journal of Paleontology. 86 (3): 488–507. doi:10.1666/11-102.1.

[CladidCrinoidsJP-73] Thomas W. Kammer; Edgar W. Roeser (2012). "Cladid Crinoids from the Late Kinderhookian Meadville Shale, Cuyahoga Formation of Ohio". Journal of Paleontology. 86 (3): 470–487. doi:10.1666/11-101.1.

[74] William I. Ausich; Mark A. Wilson (2012). "New Tethyan Apiocrinitidae (Crinoidea, Articulata) from the Jurassic of Israel". Journal of Paleontology. 86 (6): 1051–1055. doi:10.1666/12-049R.1.

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[Saaremaacrinus-76] William I. Ausich; Mark A. Wilson; Olev Vinn (2012). "Crinoids from the Silurian of Western Estonia". Acta Palaeontologica Polonica. 57 (3): 613–631. doi:10.4202/app.2010.0094.

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[AtwoodandSumrall-78] James W. Atwood and Colin D. Sumrall (2012). "Morphometric Investigation of the Pentremites Fauna from the Glen Dean Formation, Kentucky". Journal of Paleontology. 86 (5): 813–828. doi:10.1666/12-003.1.CS1 maint: Uses authors parameter (link)

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[80] Alexander B. Doweld (2012). "Heckerocrinus, a new substitute generic name for fossil crinoid Bockia Hecker 1940 (Echinodermata: Crinoidea) non Reisinger 1924 (Vermes: Turbellaria)". Paläontologische Zeitschrift. 86 (4): 457–458. doi:10.1007/s12542-012-0148-y.

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[84] Paul W. Hearn; Bradley Deline (2012). "A new species of Paradiabolocrinus from the Upper Ordovician of central Kentucky, USA". Southeastern Geology. 49 (1): 25–30.

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[87] Stephen K. Donovan; David A. T. Harper; Zhan Renbin; Lars Stemmerik; Liu Jianbo; Svend Stouge (2012). "A primitive cladid crinoid from the Jiacun Group, Tibet (Darriwilian, Middle Ordovician)". Geological Journal. 47 (6): 653–660. doi:10.1002/gj.2436.

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[89] Louis G. Zachos (2012). "A new genus and species of echinoid (Echinoidea, Spatangoida) from the Oligocene (Rupelian) of Mississippi". Southeastern Geology. 49 (1): 43–48.

[90] Junfeng Guo; Yong Li; Huiping Han; Qiang Ou; Jianren Zhou; Yajuan Zheng (2012). "New Macroscopic Problematic Fossil from the Early Cambrian Yanjiahe Biota, Yichang, Hubei, China". Acta Geologica Sinica (English Edition). 86 (4): 791–798. doi:10.1111/j.1755-6724.2012.00706.x.

[91] M. A. Fedonkin; P. Vickers-Rich; B. J. Swalla; P. Trusler; M. Hall (2012). "A new metazoan from the Vendian of the White Sea, Russia, with possible affinities to the ascidians". Paleontological Journal. 46 (1): 1–11. doi:10.1134/S0031030112010042.

[MazzaCauRigo-92] Michele Mazza; Andrea Cau; Manuel Rigo (2012). "Application of numerical cladistic analyses to the Carnian–Norian conodonts: a new approach for phylogenetic interpretations". Journal of Systematic Palaeontology. 10 (3): 401–422. doi:10.1080/14772019.2011.573584.

[NMMNHSB64-93] Michael J. Orchard (2014). "Conodonts from the Carnian-Norian Boundary (Upper Triassic) of Black Bear Ridge, Northeastern British Columbia, Canada". New Mexico Museum of Natural History and Science Bulletin. 64: 1–139.

[Mazzaetal-94] Michele Mazza; Manuel Rigo; Maria Gullo (2012). "Taxonomy and biostratigraphic record of the Upper Triassic conodonts of the Pizzo Mondello section (western Sicily, Italy), GSSP candidate for the base of the Norian". Rivista Italiana di Paleontologia e Stratigrafia. 118 (1): 85–130.

[DrygantSzaniawskiAPPconodonts-95] Daniel Drygant; Hubert Szaniawski (2012). "Lochkovian conodonts from Podolia, Ukraine, and their stratigraphic significance". Acta Palaeontologica Polonica. 57 (4): 833–861. doi:10.4202/app.2012.0124.

[TdGconodonts-96] C. A. Méndez (2012). "Moscovian conodonts from the Nevandi River Valley (Cantabrian Zone, North Spain)". Trabajos de Geología. 32: 63–75.

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[Stouge2012-100] Svend Stouge (2012). "Middle Ordovician (late Dapingian–Darriwilian) conodonts from the Cow Head Group and Lower Head Formation, western Newfoundland, Canada". Canadian Journal of Earth Sciences. 49 (1): 59–90. doi:10.1139/e11-057.

[101] Viktor Karádi; Heinz W. Kozur; Ágnes Görögt (2013). "Stratigraphically important lower Norian conodonts from the Csővár borehole (CSV-1), Hungary – Comparison with the conodont succession of the Norian GSSP candidate Pizzo Mondello" (PDF). New Mexico Museum of Natural History and Science Bulletin. 61: 284–295.

[102] Liming Gong; Chengyuan Wang; Changsheng Wang; Yunxi Hu; Yongyang Tang; Ping Hu; Qingdong Wang (2012). "Devonian conodonts in SE area of Chongqing City and its geological significance". Acta Micropalaeontologica Sinica. 29 (3): 282–298.

[103] Harald Tragelehn; Sven Hartenfels (2012). "Pseudopolygnathus inordinatus nov. nom. – homonym replacement for an upper Famennian (Upper Devonian) conodont species" (PDF). Subcommission on Devonian Stratigraphy Newsletter. 27: 59–60.

[104] Hillary C. Maddin; Farish A. Jenkins Jr; Jason S. Anderson (2012). "The Braincase of Eocaecilia micropodia (Lissamphibia, Gymnophiona) and the Origin of Caecilians". PLoS ONE. 7 (12): e50743. doi:10.1371/journal.pone.0050743. PMC 3515621. PMID 23227204.

[105] Marylène Danto; Florian Witzmann; Johannes Müller (2012). "Redescription and phylogenetic relationships of Solenodonsaurus janenschi Broili, 1924, from the Late Carboniferous of Nýřany, Czech Republic". Fossil Record. 15 (2): 45–59. doi:10.1002/mmng.201200003.

[106] Stephanie E. Pierce; Jennifer A. Clack; John R. Hutchinson (2012). "Three-dimensional limb joint mobility in the early tetrapod Ichthyostega". Nature. 486 (7404): 523–526. doi:10.1038/nature11124. PMID 22722854.

[Ymeria-107] Clack, J.A.; Ahlberg, P.E.; Blom, H.; Finney, S.M. (2012). "A new genus of Devonian tetrapod from North-East Greenland, with new information on the lower jaw of Ichthyostega". Palaeontology. 55 (1): 73–86. doi:10.1111/j.1475-4983.2011.01117.x.

[Arachana-108] Graciela Piñeiro, Alejandro Ramos and Claudia Marsicano (2012). "A rhinesuchid-like temnospondyl from the Permo-Triassic of Uruguay". Comptes Rendus Palevol. 11 (1): 65–78. doi:10.1016/j.crpv.2011.07.007.CS1 maint: Uses authors parameter (link)

[Benthosuchusgusevae-109] I. V. Novikov (2012). "New data on trematosauroid labyrinthodonts of Eastern Europe: 4. Genus Benthosuchus Efremov, 1937". Paleontological Journal. 46 (4): 400–411. doi:10.1134/S0031030112040089.CS1 maint: Uses authors parameter (link)

[Cacopswoehri-110] Fröbisch, N.B.; and Reisz, R.R. (2012). "A new species of dissorophid (Cacops woehri) from the Lower Permian Dolese Quarry, near Richards Spur, Oklahoma". Journal of Vertebrate Paleontology. 32 (1): 35–44. doi:10.1080/02724634.2012.633586.

[111] Anne Warren (2012). "The South African stereospondyl Microposaurus from the Middle Triassic of the Sydney Basin, Australia". Journal of Vertebrate Paleontology. 32 (3): 538–544. doi:10.1080/02724634.2012.658934.

[Nyranerpeton-112] Ralf Werneburg (2012). "Dissorophoide Amphibien aus dem Westphalian D (Ober-Karbon) von Nýřany in Böhmen (Tschechische Republik) - der Schlüssel zum Verständnis der frühen 'Branchiosaurier'". Semana. Naturwissenschaftliche Veröffentlichungen des Naturhistorischen Museums Schloss Bertholdsburg Schleusingen. 27: 3–50.CS1 maint: Uses authors parameter (link)

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[115] Jozef Klembara; J. Sébastien Steyer (2012). "A new species of Sclerocephalus (Temnospondyli: Stereospondylomorpha) from the Early Permian of the Boskovice Basin (Czech Republic)". Journal of Paleontology. 86 (2): 302–310. doi:10.1666/11-051.1.

[Altenglanerpeton-116] Glienke, S. (2012). "A new "microsaur" (Amphibia; Lepospondyli) from the Rotliegend of the Saar–Palatinate region (Carboniferous/Permian transition; West Germany)". Paläontologische Zeitschrift. 86 (3): 297–311. doi:10.1007/s12542-012-0130-8.

[117] Márton Venczel; Vlad Codrea; Cristina Fărcaş (2013). "A new palaeobatrachid frog from the early Oligocene of Suceag, Romania". Journal of Systematic Palaeontology. 11 (2): 179–189. doi:10.1080/14772019.2012.671790.

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[120] Ke-Qin Gao; Neil H. Shubin (2012). "Late Jurassic salamandroid from western Liaoning, China". Proceedings of the National Academy of Sciences of the United States of America. 109 (15): 5767–5772. doi:10.1073/pnas.1009828109. PMC 3326464. PMID 22411790.

[SGS482Calyptocephalellidae-121] Federico Agnolin (2012). "A new Calyptocephalellidae (Anura, Neobatrachia) from the Upper Cretaceous of Patagonia, Argentina, with comments on its systematic position". Studia Geologica Salmanticensia. 48 (2): 129–178.

[122] James D. Gardner (2012). "Revision of Piceoerpeton Meszoely (Caudata: Scapherpetontidae) and description of a new species from the late Maastrichtian and ?early Paleocene of western North America". Bulletin de la Société Géologique de France. 183 (6): 611–620. doi:10.2113/gssgfbull.183.6.611.CS1 maint: Uses authors parameter (link)

[Graellsia681-123] C. Martín; M. A. Alonso-Zarazaga; B. Sanchiz (2012). "Nomenclatural notes on living and fossil amphibians". Graellsia. 68 (1): 159–180. doi:10.3989/graellsia.2012.v68.056.

[124] Fabien Laloy; Jean-Claude Rage; Susan E. Evans; Renaud Boistel; Nicolas Lenoir; Michel Laurin (2013). "A re-interpretation of the Eocene anuran Thaumastosaurus based on microCT examination of a 'mummified' specimen". PLoS ONE. 8 (9): e74874. doi:10.1371/journal.pone.0074874. PMC 3783478. PMID 24086389.

[Seminobatrachus-125] Pavel P. Skutschas; Yuri M. Gubin (2012). "A new salamander from the late Paleocene–early Eocene of Ukraine". Acta Palaeontologica Polonica. 57 (1): 135–148. doi:10.4202/app.2010.0101.

[126] Ana M. Báez; Raúl O. Gómez; Luiz C.B. Ribeiro; Agustín G. Martinelli; Vicente P.A. Teixeira; Mara L.F. Ferraz (2012). "The diverse Cretaceous neobatrachian fauna of South America: Uberabatrachus carvalhoi, a new frog from the Maastrichtian Marília Formation, Minas Gerais, Brazil". Gondwana Research. 22 (3–4): 1141–1150. doi:10.1016/j.gr.2012.02.021.

[127] Jocelyn Falconnet (2012). "First evidence of a bolosaurid parareptile in France (latest Carboniferous-earliest Permian of the Autun basin) and the spatiotemporal distribution of the Bolosauridae". Bulletin de la Société Géologique de France. 183 (6): 495–508. doi:10.2113/gssgfbull.183.6.495.CS1 maint: Uses authors parameter (link)

[128] Mark J. Macdougall and Robert Reisz (2012). "A new parareptile (Parareptilia, Lanthanosuchoidea) from the Early Permian of Oklahoma". Journal of Vertebrate Paleontology. 32 (5): 1018–1026. doi:10.1080/02724634.2012.679757.CS1 maint: Uses authors parameter (link)

[129] Jocelyn Falconnet, Misalalanirina Andriamihaja, Émilie Läng and J.-Sébastien Steyer (2012). "First procolophonid (Reptilia, Parareptilia) from the Lower Triassic of Madagascar". Comptes Rendus Palevol. 11 (5): 357–369. doi:10.1016/j.crpv.2012.04.001.CS1 maint: Uses authors parameter (link)

[Acamptonectes-130] Fischer,V.; Maisch, M.W.; Naish, D.; Kosma, R.; Liston, J.; Joger, U.; Krüger, F.J.; Pérez,J.P.; Tainsh, J.; Appleby, R.M. (2012). "New Ophthalmosaurid Ichthyosaurs from the European Lower Cretaceous Demonstrate Extensive Ichthyosaur Survival across the Jurassic–Cretaceous Boundary". PLoS ONE. 7 (1): e29234. doi:10.1371/journal.pone.0029234. PMC 3250416. PMID 22235274.CS1 maint: Uses authors parameter (link)

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[132] Erin E. Maxwell, Marta S. Fernández and Rainer R. Schoch (2012). "First Diagnostic Marine Reptile Remains from the Aalenian (Middle Jurassic): A New Ichthyosaur from Southwestern Germany". PLoS ONE. 7 (8): e41692. doi:10.1371/journal.pone.0041692. PMC 3411580. PMID 22870244.CS1 maint: Uses authors parameter (link)

[133] Jeremy E. Martin; Valentin Fischer; Peggy Vincent; Guillaume Suan (2012). "A longirostrine Temnodontosaurus (Ichthyosauria) with comments on Early Jurassic ichthyosaur niche partitioning and disparity". Palaeontology. 55 (5): 995–1005. doi:10.1111/j.1475-4983.2012.01159.x.

[Largocephalosaurus-134] Long Cheng, Xiaohong Chen, Xiongwei Zeng and Yongjian Cai (2012). "A new eosauropterygian (Diapsida: Sauropterygia) from the Middle Triassic of Luoping, Yunnan Province". Journal of Earth Science. 23 (1): 33–40. doi:10.1007/s12583-012-0231-z.CS1 maint: Uses authors parameter (link)

[135] Chun Li, Da-Yong Jiang, Long Cheng, Xiao-Chun Wu and Olivier Rieppel (2014). "A new species of Largocephalosaurus (Diapsida: Saurosphargidae), with implications for the morphological diversity and phylogeny of the group". Geological Magazine. 151 (1): 100–120. doi:10.1017/S001675681300023X.CS1 maint: Uses authors parameter (link)

[136] Tai Kubo; Mark T. Mitchell; Donald M. Henderson (2012). "Albertonectes vanderveldei, a new elasmosaur (Reptilia, Sauropterygia) from the Upper Cretaceous of Alberta". Journal of Vertebrate Paleontology. 32 (3): 557–572. doi:10.1080/02724634.2012.658124.

[137] Peggy Vincent and Roger B. J. Benson (2012). "Anningasaura, a basal plesiosaurian (Reptilia, Plesiosauria) from the Lower Jurassic of Lyme Regis, United Kingdom". Journal of Vertebrate Paleontology. 32 (5): 1049–1063. doi:10.1080/02724634.2012.686467.CS1 maint: Uses authors parameter (link)

[PLOSTriassicJurassicBoundary-138] Roger B. J. Benson, Mark Evans and Patrick S. Druckenmiller (2012). "High Diversity, Low Disparity and Small Body Size in Plesiosaurs (Reptilia, Sauropterygia) from the Triassic–Jurassic Boundary". PLoS ONE. 7 (3): e31838. doi:10.1371/journal.pone.0031838. PMC 3306369. PMID 22438869.CS1 maint: Uses authors parameter (link)

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[D.tropicensis-140] Rebecca Schmeisser McKean (2012). "A new species of polycotylid plesiosaur (Reptilia: Sauropterygia) from the Lower Turonian of Utah: extending the stratigraphic range of Dolichorhynchops". Cretaceous Research. 34: 184–199. doi:10.1016/j.cretres.2011.10.017.CS1 maint: Uses authors parameter (link)

[Vectocleidus-141] Roger B. J. Benson, Hilary F. Ketchum, Darren Naish and Langan E. Turner (2013). "A new leptocleidid (Sauropterygia, Plesiosauria) from the Vectis Formation (Early Barremian–early Aptian; Early Cretaceous) of the Isle of Wight and the evolution of Leptocleididae, a controversial clade". Journal of Systematic Palaeontology. 11 (2): 233–250. doi:10.1080/14772019.2011.634444.CS1 maint: Uses authors parameter (link)

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[144] Espen M. Knutsen, Patrick S. Druckenmiller and Jørn H. Hurum (2012). "A new species of Pliosaurus (Sauropterygia: Plesiosauria) from the Middle Volgian of central Spitsbergen, Norway". Norwegian Journal of Geology. 92 (2–3): 235–258.CS1 maint: Uses authors parameter (link)

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[147] Oliver W. M. Rauhut; Alexander M. Heyng; Adriana López-Arbarello; Andreas Hecker (2012). "A New Rhynchocephalian from the Late Jurassic of Germany with a Dentition That Is Unique amongst Tetrapods". PLoS ONE. 7 (10): e46839. doi:10.1371/journal.pone.0046839. PMC 3485277. PMID 23118861.

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[arretosaurids-152] V. R. Alifanov (2012). "Lizards of the family Arretosauridae Gilmore, 1943 (Iguanomorpha, Iguania) from the Paleogene of Mongolia". Paleontological Journal. 46 (4): 412–420. doi:10.1134/S0031030112040028.

[Eremiasaurus-153] Leblanc, A.R.H.; Caldwell, M.W.; Bardet, N. (2012). "A new mosasaurine from the Maastrichtian (Upper Cretaceous) phosphates of Morocco and its implications for mosasaurine systematics". Journal of Vertebrate Paleontology. 32 (1): 82–104. doi:10.1080/02724634.2012.624145.

[154] Boler, Arnau; Susan E. Evans (2012). "A tiny lizard (Lepidosauria, Squamata) from the Lower Cretaceous of Spain". Palaeontology. 55 (3): 491–500. doi:10.1111/j.1475-4983.2012.01145.x.

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[156] László Makádi; Michael W. Caldwell; Attila Ősi (2012). "The First Freshwater Mosasauroid (Upper Cretaceous, Hungary) and a New Clade of Basal Mosasauroids". PLoS ONE. 7 (12): e51781. doi:10.1371/journal.pone.0051781. PMC 3526648. PMID 23284766.

[157] Robert M. Sullivan, Marc Augé, Eric Wille and Richard Smith (2012). "A new glyptosaurine lizard from the earliest Eocene of Dormaal, Belgium". Bulletin de la Société Géologique de France. 183 (6): 627–633. doi:10.2113/gssgfbull.183.6.627.CS1 maint: Uses authors parameter (link)

[AlifanovLacertilia-158] V. R. Alifanov (2012). "ОТРЯД LACERTILIA". In E.N. Kurochkin; A. V. Lopatin. Ископаемые позвоночные России и сопредельных стран. Ископаемые рептилии и птицы. Часть 2 / Fossil vertebrates of Russia and adjacent countries. Fossil Reptiles and Birds. Part 2. GEOS. pp. 7–136. ISBN 978-5-89118-594-4.

[159] Jozef Klembara (2012). "A new species of Pseudopus (Squamata, Anguidae) from the early Miocene of Northwest Bohemia (Czech Republic)". Journal of Vertebrate Paleontology. 32 (4): 854–866. doi:10.1080/02724634.2012.670177.

[160] Susan E. Evans, Marc E. H. Jones and Ryoko Matsumoto (2012). "A new lizard skull from the Purbeck Limestone Group (Lower Cretaceous) of England". Bulletin de la Société Géologique de France. 183 (6): 517–524. doi:10.2113/gssgfbull.183.6.517.CS1 maint: Uses authors parameter (link)

[161] Salvador Bailon and Marc Auge (2012). "A news genus, Ragesaurus (Squamata, Anguidae, Anguinae) from the lower Pleistocene of the islas Medas (Catalonia, Spain)". Bulletin de la Société Géologique de France. 183 (6): 683–688. doi:10.2113/gssgfbull.183.6.683.CS1 maint: Uses authors parameter (link)

[162] Juan D. Daza; Aaron M. Bauer (2012). "A new amber-embedded sphaerodactyl gecko from Hispaniola, with comments on morphological synapomorphies of the Sphaerodactylidae" (PDF). Breviora. 529: 1–28. doi:10.3099/529.1.

[163] Juan D. Daza; Virginia Abdala; J. Salvador Arias; Daniel García-López; Pablo Ortiz (2012). "Cladistic Analysis of Iguania and a Fossil Lizard from the Late Pliocene of Northwestern Argentina". Journal of Herpetology. 46 (1): 104–119. doi:10.1670/10-112.

[164] Jack L. Conrad; Ana M. Balcarcel; Carl M. Mehling (2012). "Earliest Example of a Giant Monitor Lizard (Varanus, Varanidae, Squamata)". PLoS ONE. 7 (8): e41767. doi:10.1371/journal.pone.0041767. PMC 3416840. PMID 22900001.

[165] Andrea Villa; Juan Abella; David M. Alba; Sergio Almécija; Arnau Bolet; George D. Koufos; Fabien Knoll; Àngel H. Luján; Jorge Morales; Josep M. Robles; Israel M. Sánchez; Massimo Delfino (2018). "Revision of Varanus marathonensis (Squamata, Varanidae) based on historical and new material: morphology, systematics, and paleobiogeography of the European monitor lizards". PLoS ONE. 13 (12): e0207719. doi:10.1371/journal.pone.0207719. PMC 6281198. PMID 30517172.

[AnquetinJSP2012-166] Jérémy Anquetin (2012). "Reassessment of the phylogenetic interrelationships of basal turtles (Testudinata)". Journal of Systematic Palaeontology. 10 (1): 3–45. doi:10.1080/14772019.2011.558928.

[167] Hans-Volker Karl, Elke Gröning and Carsten Brauckmann (2012). "New materials of the giant sea turtle Allopleuron (Testudines: Chelonioidea) from the marine Late Cretaceous of Central Europe and the Palaeogene of Kazakhstan". Stvdia Palaeocheloniologica IV, Stvdia Geologica Salmanticensia. n Especial 9: 153–173.CS1 maint: Uses authors parameter (link)

[Vitek-168] Natasha S. Vitek (2012). "Giant fossil soft-shelled turtles of North America". Palaeontologia Electronica. 15 (1): 13A.CS1 maint: Uses authors parameter (link)

[169] Hans-Volker Karl, Elke Gröning, Carsten Brauckmann and Mike Reich (2012). "Ballerstedtia bueckebergensis, a new turtle from the Early Cretaceous Wealden Facies of Germany (Testudines: Pleurosternidae)". Stvdia Palaeocheloniologica IV, Stvdia Geologica Salmanticensia. n Especial 9: 47–60.CS1 maint: Uses authors parameter (link)

[Sichuanturtles-170] Haiyan Tong, Igor Danilov, Yong Ye, Hui Ouyang and Guangzhao Peng (2012). "Middle Jurassic turtles from the Sichuan Basin, China: a review". Geological Magazine. 149 (4): 675–695. doi:10.1017/S0016756811000859.CS1 maint: Uses authors parameter (link)

[171] Adán Pérez-García (2012). "Berruchelus russelli, gen. et sp. nov., a paracryptodiran turtle from the Cenozoic of Europe". Journal of Vertebrate Paleontology. 32 (3): 545–556. doi:10.1080/02724634.2012.658933.

[172] Edwin A. Cadena; Daniel T. Ksepka; Carlos A. Jaramillo; Jonathan I. Bloch (2012). "New pelomedusoid turtles from the late Palaeocene Cerrejón Formation of Colombia and their implications for phylogeny and body size evolution". Journal of Systematic Palaeontology. 10 (2): 313–331. doi:10.1080/14772019.2011.569031.

[173] Márton Rabi, Haiyan Tong and Gábor Botfalvai (2012). "A new species of the side-necked turtle Foxemys (Pelomedusoides: Bothremydidae) from the Late Cretaceous of Hungary and the historical biogeography of the Bothremydini". Geological Magazine. 149 (4): 662–674. doi:10.1017/S0016756811000756.CS1 maint: Uses authors parameter (link)

[174] Adán Pérez-García; Xabier Murelaga (2012). "Galvechelone lopezmartinezae gen. et sp. nov., a new cryptodiran turtle in the Lower Cretaceous of Europe". Palaeontology. 55 (5): 937–944. doi:10.1111/j.1475-4983.2012.01154.x.

[175] Julien Claude, Jiang-Yong Zhang, Jian-Jun Li, Jin-You Mo, Xue-Wen Kuang and Haiyan Tong (2012). "Geoemydid turtles from the Late Eocene Maoming basin, southern China". Bulletin de la Société Géologique de France. 183 (6): 641–651. doi:10.2113/gssgfbull.183.6.641.CS1 maint: Uses authors parameter (link)

[Hoyasemys-176] Adán Pérez-García, Marcelo S. de la Fuente and Francisco Ortega (2012). "Hoyasemys jimenezi gen. et sp. nov., a freshwater basal eucryptodiran turtle from the Lower Cretaceous of Spain". Acta Palaeontologica Polonica. 57 (2): 285–298. doi:10.4202/app.2011.0031.CS1 maint: Uses authors parameter (link)

[177] Adán Pérez-García, Francisco Ortega, Xabier Murelaga (2012). "A new genus of Bothremydidae (Chelonii, Pleurodira) in the Cretaceous of Southwestern Europe". Geobios. 45 (2): 219–229. doi:10.1016/j.geobios.2011.03.001.CS1 maint: Uses authors parameter (link)

[178] Jason R. Bourque (2012). "An extinct mud turtle of the Kinosternon flavescens group (Testudines, Kinosternidae) from the middle Miocene (late Barstovian) of New Mexico". Journal of Vertebrate Paleontology. 32 (1): 68–81. doi:10.1080/02724634.2012.626824.CS1 maint: Uses authors parameter (link)

[179] Jason R. Bourque (2012). "A fossil mud turtle (Testudines, Kinosternidae) from the early middle Miocene (early Barstovian) of New Mexico". Journal of Vertebrate Paleontology. 32 (4): 836–853. doi:10.1080/02724634.2012.670179.CS1 maint: Uses authors parameter (link)

[180] Adán Pérez-García and Xabier Murelaga (2012). "Larachelus morla, gen. et sp. nov., a new member of the little-known European Early Cretaceous record of stem cryptodiran turtles". Journal of Vertebrate Paleontology. 32 (6): 1293–1302. doi:10.1080/02724634.2012.694592.CS1 maint: Uses authors parameter (link)

[181] Hans-Volker Karl, Stefanie Biermann and Gottfried Tichy (2012). "The oldest true sea turtle of the world, Oertelia gigantea (Oertel, 1914) n. gen. from the Aptian of Kastendamm near Hanover, Germany". Stvdia Palaeocheloniologica IV, Stvdia Geologica Salmanticensia. n Especial 9: 107–128.CS1 maint: Uses authors parameter (link)

[182] Juliana Sterli; Marcelo S. de la Fuente (2013). "New evidence from the Palaeocene of Patagonia (Argentina) on the evolution and palaeo-biogeography of Meiolaniformes (Testudinata, new taxon name)". Journal of Systematic Palaeontology. 11 (7): 835–852. doi:10.1080/14772019.2012.708674.

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[184] J. Marmi; Á.H. Luján; V. Riera; R. Gaete; O. Oms; À. Galobart (2012). "The youngest species of Polysternon: A new bothremydid turtle from the uppermost Maastrichtian of the southern Pyrenees". Cretaceous Research. 35: 133–142. doi:10.1016/j.cretres.2011.12.004.

[185] Edwin A. Cadena; Jonathan I. Bloch; Carlos A. Jaramillo (2012). "New Bothremydid Turtle (Testudines, Pleurodira) from the Paleocene of Northeastern Colombia". Journal of Paleontology. 86 (4): 688–698. doi:10.1666/11-128R1.1.

[Cadenaetal-186] Edwin Cadena; Jason R. Bourque; Aldo F. Rincon; Jonathan I. Bloch; Carlos A. Jaramillo; Bruce J. Macfadden (2012). "New Turtles (Chelonia) from the Late Eocene Through Late Miocene of the Panama Canal Basin". Journal of Paleontology. 86 (3): 539–557. doi:10.1666/11-106.1.

[187] Hans-Volker Karl and Henrik Madsen (2012). "Tasbacka danica n. sp., a new Eocene marine turtle of Denmark (Testudines: Chelonioidea)". Stvdia Palaeocheloniologica IV, Stvdia Geologica Salmanticensia. n Especial 9: 193–204.CS1 maint: Uses authors parameter (link)

[188] Walter G. Joyce, Andrea Petričević, Tyler R. Lyson and Nicholas J. Czaplewski (2012). "A New Box Turtle from the Miocene/Pliocene Boundary (Latest Hemphillian) of Oklahoma and A Refined Chronology of Box Turtle Diversification". Journal of Paleontology. 86 (1): 177–190. doi:10.1666/11-073.1.CS1 maint: Uses authors parameter (link)

[189] Haiyan Tong, Li Xu, Eric Buffetaut, Xingliao Zhang and Songhai Jia (2012). "A new nanhsiungchelyid turtle from the Late Cretaceous of Neixiang, Henan Province, China". Annales de Paléontologie. 98 (4): 303–314. doi:10.1016/j.annpal.2012.08.001.CS1 maint: Uses authors parameter (link)

[JVPCischigualastensis-190] María J. Trotteyn; Ricardo N. Martínez; Oscar A. Alcober (2012). "A new proterochampsid Chanaresuchus ischigualastensis (Diapsida, Archosauriformes) in the early Late Triassic Ischigualasto Formation, Argentina". Journal of Vertebrate Paleontology. 32 (2): 485–489. doi:10.1080/02724634.2012.645975.

[191] María Jimena Trotteyn; Martín D. Ezcurra (2014). "Osteology of Pseudochampsa ischigualastensis gen. et comb. nov. (Archosauriformes: Proterochampsidae) from the Early Late Triassic Ischigualasto Formation of Northwestern Argentina". PLoS ONE. 9 (11): e111388. doi:10.1371/journal.pone.0111388. PMC 4245112. PMID 25426846.

[Diandongosuchus-192] Chun Li; Xiao-Chun Wu; Li-Jun Zhao; Tamaki Sato & Li-Ting Wang (2012). "A new archosaur (Diapsida, Archosauriformes) from the marine Triassic of China". Journal of Vertebrate Paleontology. 32 (5): 1064–1081. doi:10.1080/02724634.2012.694383.

[193] Michelle R. Stocker; Sterling J. Nesbitt; Li-Jun Zhao; Xiao-Chun Wu; Chun Li (2016). "Mosaic evolution in Phytosauria: the origin of long-snouted morphologies based on a complete skeleton of a phytosaur from the Middle Triassic of China". Society of Vertebrate Paleontology 76th Annual Meeting Program & Abstracts: 232.

[194] Michelle R. Stocker; Li-Jun Zhao; Sterling J. Nesbitt; Xiao-Chun Wu; Chun Li (2017). "A Short-Snouted, Middle Triassic Phytosaur and its Implications for the Morphological Evolution and Biogeography of Phytosauria". Scientific Reports. 7: Article number 46028. doi:10.1038/srep46028.

[195] Andrew B. Heckert; Spencer G. Lucas; Justin A. Spielmann (2012). "A new species of the enigmatic archosauromorph Doswellia from the Upper Triassic Bluewater Creek Formation, New Mexico, USA". Palaeontology. 55 (6): 1333–1348. doi:10.1111/j.1475-4983.2012.01200.x.

[196] Michelle R. Stocker (2012). "A new phytosaur (Archosauriformes, Phytosauria) from the Lot's Wife beds (Sonsela Member) within the Chinle Formation (Upper Triassic) of Petrified Forest National Park, Arizona". Journal of Vertebrate Paleontology. 32 (3): 573–586. doi:10.1080/02724634.2012.649815.

[197] Roger B. J. Benson (2012). "Interrelationships of basal synapsids: cranial and postcranial morphological partitions suggest different topologies". Journal of Systematic Palaeontology. 10 (4): 601–624. doi:10.1080/14772019.2011.631042.CS1 maint: Uses authors parameter (link)

[PJKurkin-198] A. A. Kurkin (2012). "Dicynodontids of Eastern Europe". Paleontological Journal. 46 (2): 187–198. doi:10.1134/S003103011201008X.CS1 maint: Uses authors parameter (link)

[KAF11-199] Kammerer, C.F.; Angielczyk, K.D.; Fröbisch, J. (2011). "A comprehensive taxonomic revision of Dicynodon (Therapsida, Anomodontia) and its implications for dicynodont phylogeny, biogeography, and biostratigraphy". Journal of Vertebrate Paleontology. 31 (Suppl. 1): 1–158. doi:10.1080/02724634.2011.627074.

[PJIvakhnenko-200] M. F. Ivakhnenko (2012). "Permian Cynodontia (Theromorpha) of Eastern Europe". Paleontological Journal. 46 (2): 199–207. doi:10.1134/S0031030112020062.

[201] Juan Carlos Cisneros, Fernando Abdala, Saniye Atayman-Güven, Bruce S. Rubidge, A. M. Celâl Şengör and Cesar L. Schultz (2012). "Carnivorous dinocephalian from the Middle Permian of Brazil and tetrapod dispersal in Pangaea". Proceedings of the National Academy of Sciences of the United States of America. 109 (5): 1584–1588. doi:10.1073/pnas.1115975109. PMC 3277192. PMID 22307615.CS1 maint: Uses authors parameter (link)

[202] D. P. Das and Abir Gupta (2012). "New cynodont record from the lower Triassic Panchet Formation, Damodar valley". Journal of the Geological Society of India. 79 (2): 175–180. doi:10.1007/s12594-012-0022-2.CS1 maint: Uses authors parameter (link)

[Purbeckodon-203] P. M. Butler, D. Sigogneau-Russell and P. C. Ensom (2012). "Possible persistence of the morganucodontans in the Lower Cretaceous Purbeck Limestone Group (Dorset, England)". Cretaceous Research. 33 (1): 135–145. doi:10.1016/j.cretres.2011.09.007.CS1 maint: Uses authors parameter (link)

[Aggerscolex-204] Joseph P. Botting; Lucy A. Muir; Peter Van Roy; Denis Bates; Christopher Upton (2012). "Diverse middle Ordovician palaeoscolecidan worms from the Builth-Llandrindod Inlier of central Wales". Palaeontology. 55 (3): 501–528. doi:10.1111/j.1475-4983.2012.01135.x.

[205] Joachim T. Haug; Georg Mayer; Carolin Haug; Derek E.G. Briggs (2012). "A Carboniferous Non-Onychophoran Lobopodian Reveals Long-Term Survival of a Cambrian Morphotype". Current Biology. 22 (18): 1673–1675. doi:10.1016/j.cub.2012.06.066. PMID 22885062.

[HyolithaBG-206] Martin Valent; Oldřich Fatka; Michal Szabad; Václav Micka; Ladislav Marek (2012). "Two new orthothecids from the Cambrian of the Barrandian area (Hyolitha, Skryje-Týřovice Basin, Czech Republic)". Bulletin of Geosciences. 87 (2): 241–248. doi:10.3140/bull.geosci.1142.

[207] Erica C. Clites; Mary L. Droser; James G. Gehling (2012). "The advent of hard-part structural support among the Ediacara biota: Ediacaran harbinger of a Cambrian mode of body construction". Geology. 40 (4): 307–310. doi:10.1130/G32828.1.

[208] Shuji Niko (2012). "Additional Material of Favositid and Auloporid Tabulate Corals from the Devonian Fukuji Formation, Gifu Prefecture, Japan" (PDF). Bulletin of the National Museum of Nature and Science, Series C. 38: 47–62.

[Dispinoscolex-209] Baichuan Duan; Xi-Ping Dong; Philip C. J. Donoghue (2012). "New palaeoscolecid worms from the Furongian (upper Cambrian) of Hunan, South China: is Markuelia an embryonic palaeoscolecid?". Palaeontology. 55 (3): 613–622. doi:10.1111/j.1475-4983.2012.01148.x.

[BAP383-210] Ramón Andrés López-Pérez (2012). "Late Miocene to Pleistocene Reef Corals in the Gulf of California". Bulletins of American Paleontology. 383: 1–78.

[Nematology144-211] George O. Poinar Jr (2012). "New fossil nematodes in Dominican and Baltic amber". Nematology. 14 (4): 483–488. doi:10.1163/156854111X612199.

[NikoHaikawaFujikawa-212] Shuji Niko; Takehiko Haikawa; Masayuki Fujikawa (2012). "Permian tabulate corals from the Akiyoshi Limestone Group, Yamaguchi Prefecture". Bulletin of the Akiyoshi-dai Museum of Natural History. 47: 5–10.

[Redondagnathus-213] Justin A. Spielmann; Spencer G. Lucas (2012). "Tetrapod fauna of the Upper Triassic Redonda Formation, east-central New Mexico; the characteristic assemblage of the Apachean land-vertebrate faunachron". New Mexico Museum of Natural History and Science Bulletin. 55: 1–119.

[214] Spencer G. Lucas; John W. Estep; Andrew B. Heckert; Adrian P. Hunt (1999). "Cynodont teeth from the Upper Triassic of New Mexico, USA". Neues Jahrbuch für Geologie und Paläontologie - Monatshefte. Jg. 1999 (6): 331–344.

[215] F. Abdala; J. Neveling; J. Welman (2006). "A new trirachodontid cynodont from the lower levels of the Burgersdorp Formation (Lower Triassic) of the Beaufort Group, South Africa and the cladistic relationships of Gondwanan gomphodonts". Zoological Journal of the Linnean Society. 147 (3): 383–413. doi:10.1111/j.1096-3642.2006.00224.x.

[216] Christian A. Sidor; James A. Hopson (2018). "Cricodon metabolus (Cynodontia: Gomphodontia) from the Triassic Ntawere Formation of northeastern Zambia: patterns of tooth replacement and a systematic review of the Trirachodontidae". Journal of Vertebrate Paleontology. 37 (Supplement to No. 6): 39–64. doi:10.1080/02724634.2017.1410485.

[217] Andrzej Baliński; Yuanlin Sun; Jerzy Dzik (2012). "470-Million-year-old black corals from China". Naturwissenschaften. 99 (8): 645–653. doi:10.1007/s00114-012-0947-8. PMC 3407557.

[218] Lorna J. O'Brien; Jean-Bernard Caron (2012). "A New Stalked Filter-Feeder from the Middle Cambrian Burgess Shale, British Columbia, Canada". PLoS ONE. 7 (1): e29233. doi:10.1371/journal.pone.0029233. PMC 3261148. PMID 22279532.

[219] Shuji Niko; Yousuke Ibaraki (2012). "Syringopora konishii, a new species of Early Carboniferous tabulate corals from the allochthonous limestone blocks in the Kotaki Formation, Niigata Prefecture, central Japan". Science reports of Niigata University. (Geology). 27: 9–14.

[220] Shuji Niko; Tomio Adachi (2012). "Silurian Favositids (Coelenterata: Tabulata) from the Gionyama Formation, Miyazaki Prefecture, Japan" (PDF). Bulletin of the National Museum of Nature and Science, Series C. 38: 33–46.

[221] Shuji Niko (2012). "Trachypsammia konbo, a new species of Early Carboniferous tabulate corals from the Akiyoshi Limestone Group, Yamaguchi Prefecture". Bulletin of the Akiyoshi-dai Museum of Natural History. 47: 1–4.

[222] William J. Woelkerling; Davide Bassi; Yasufumi Iryu (2012). "Hydrolithon braganum sp. nov. (Corallinaceae, Rhodophyta), the first known exclusively fossil semi-endophytic coralline red alga". Phycologia. 51 (6): 604–611. doi:10.2216/11-119.1.

[223] Michael Krings; Thomas N. Taylor; Nora Dotzler; Gianna Persichini (2012). "Fossil fungi with suggested affinities to the Endogonaceae from the Middle Triassic of Antarctica". Mycologia. 104 (4): 835–844. doi:10.3852/11-384. PMID 22453117.

2012 in paleontology

Contents

Plants

Gymnosperms

Angiosperms

Other plants

Arthropods

Bryozoans

Brachiopods

Molluscs

Echinoderms

Ascidians

Conodonts

Fishes

Amphibians

Research

New taxa

Newly named basal tetrapods

Newly named temnospondyls

Newly named lepospondyls

Newly named lissamphibians

Parareptiles

Newly named parareptiles

Ichthyopterygians

Newly named ichthyosaurs

Lepidosauromorphs

Newly named saurosphargids

Newly named sauropterygians

Newly named rhynchocephalians

Squamates

Research

New taxa

Turtles

Research

New taxa

Archosauromorphs

Newly named basal archosauromorphs

Archosaurs

Synapsids

Non-mammalian synapsids

Research

New taxa

Mammals

Other animals

Other organisms

References

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